Jabbastygnus huttorum, Kury & M., 2015

Kury, Adriano B. & M., Osvaldo Villarreal, 2015, The prickly blade mapped: establishing homologies and a chaetotaxy for macrosetae of penis ventral plate in Gonyleptoidea (Arachnida, Opiliones, Laniatores), Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 174 (1), pp. 1-46 : 32-34

JABBASTYGNUS HUTTORUM SP. NOV.

Etymology: The Hutts are a fictional alien species in the Star Wars universe.

Type data: ♂ holotype (IAvH 3000110) Colombia, Boyacá, San Pedro de Iguaque, Morro Negro, Transecto

CL 1.7 (1.8) Ti I 1.1 (1.1) AL 1.9 (1.8) Ti II 1.8 (1.7) CW 2.4 (2.3) Ti III 1.5 (1.3) AW 2.7 (2.8) Ti IV 2.0 (2.0) ID 1.4 (1.4) Fe I 1.7 (1.5) Pp Fe 2.1 (1.9) Fe II 2.5 (2.3) Pp Pa 1.2 (1.1) Fe III 2.1 (2.0)

Fe IV 2.5 (2.4)

♂ holotype and ♂ paratype (in parentheses). A, abdomen; C, carapace; Fe, femur; ID, interocular distance; L, length; Pa, patella; Pp, pedipalpus; Ti, tibia; W, width.

1, secondary forest, pitfall trap 3245 m (05°38′32.1″N, 73°28′58.7″W), 13–15.v.2003 Elvia Lucía González & Claudia Reina leg.; 1 ♂ paratype (IAvH 3000042) Colombia, Boyacá, Villa de Leyva, Santuario de Fauna y Flora Iguaque , Sitio Laguna Iguaque 3450 m (5°38′N, 73°29′W), 16.x.1998, D. Forero leg.

Description of male holotype: Measurements of body and appendages of both holotype and paratype males in Table 7.

Body ( Fig. 18A, B ): Dorsal scutum outline alpha (subrectangular with rounded sides and widened at midlength), posterior margin slightly convex. Cheliceral sockets shallow, flanked by two pairs of small triangular processes. Anterior corners of carapace each with two setiferous tubercles. Eyes placed on separate individual mounds, space between them and interocular mound densely covered with minute granules. Interocular mound elevated, bearing two acuminate spines in longitudinal row, and placed very close to anterior margin of carapace. Posterior half of carapace strongly convex. Mesotergum divided into four areas, area I divided into left and right halves, its outline distorted both by scutal groove and by projection of area II, so both halves touch each other only at a small point. Areas III and IV partially fused in the median portion. Mesotergal area I with pair of granules, areas II–IV each with a transverse row of granules accompanied by supplementary median ones. Posterior margin of scutum and free tergites I–III each with a transverse row of acuminate setiferous tubercles.

Appendages: Cheliceral bulla with two pairs of ectal tubercles, cheliceral hand kidney-like, swollen ( Fig. 17A– C ). Pedipalpal femur and patella short, the former with longitudinal ventral row of setiferous tubercles, otherwise entirely unarmed ( Fig. 18C ). Pedipalpal tibia and tarsus each with ventro-ectal and ventro-mesal row of spines, spine count tibia (v-e IiIiIiIi and v-m IIiIi), tarsus (v-e Iiiii and v-m IiIiiii) ( Fig. 18C, D ). Legs I–IV short ( Fig. 18A ). Tr II with dorso-apical tubercle. Tr III subcubic, robust, with large retro-apical tubercle ( Fig. 17A ). Fe III with pro-dorsal, pro-ventral and retroventral rows of acuminate setiferous tubercles, the two last growing distally. Fe III also with stout proximal retro-dorsal setiferous tubercle. Ti III uniformly thick, with longitudinal rows of setiferous tubercles, the retrodorsal and retro ventral stouter. Cx IV armed with strong dorsal and pro-dorsal acuminate tubercles. Tr IV with two larger dorsal tubercles amidst smaller ones, and one retro-ventral setiferous tubercle. Fe IV ( Fig. 18 E-G) with pro-ventral and retro-ventral rows of acuminate setiferous tubercles, divergent distally and one stout retro-dorsal setiferous tubercle. Ti IV with six longitudinal rows of strong setiferous tubercles, retroventral the strongest. Basal tarsomeres IV thickened ( Fig. 18H ). Tarsal claws III–IV subparallel, unpectinated, tarsi without scopulae. Penultimate and antepenultimate tarsomeres of leg IV short, wedge-shaped. Last tarsomere of leg IV moderately elongate, intermediate in shape between the short wedge of Heterostygninae and the elongate rod of Stygninae. Tarsal counts: 6(3)–6(3)/10(4)–10(3)/6–6/7–7.

Genitalia ( Fig. 19A–D ): Truncus apical inflated as a torus encircled by multiple small wrinkles dorsal and lateral plus a few thick ventral folds. Malleus hemispherical, projected mid-dorsally as a podium where the glans rests. Malleus contains MS A1–A3 forming a dorsolateral arch and B, making part of the same arch, but separated by a wide gap and inserted ventrally. Lamina parva complex-shaped, formed by: (1) a columnar base, compressed laterally bearing MS C–E; (2) a flat subquadrangular platform projected laterally into two wide lobes; and (3) a pair of erect mid-dorsal lobes. MS D1 small, dorso-lateral, inserted distally, at the junction glans/stylus; MS C1–C3 forming a transverse row just below platform; MS E1–E2 long, forming a longitudinal ventro-lateral row.

Variation: Paratype has only one spine atop the interocular mound. Tarsal counts of paratype: 5(3)– 5(3)/9(3)–8(3)/6–5/6–6.

Figures

Figure 17. Jabbastygnus huttorum sp. nov., male holotype (IAvH 3000110): A, habitus, dorsal view; B, same, lateral view; C, carapace, pedipalpi and chelicerae, frontal view; D, free tergites and sternites, posterior view. Scale bars = 1 mm.

Figure 18. Jabbastygnus huttorum sp. nov., male holotype (IAvH 3000110): A, habitus, dorsal view; B, same, lateral view; C, right pedipalpus, ectal view; D, right pedipalpus Pa–Ta, mesal view; E, right leg IV, prolateral view; F, same, dorsal view; G, same, ventral view; H, tarsus IV, lateral view. Scale bars = 1 mm.

Figure 19. Jabbastygnus huttorum sp. nov., male holotype (IAvH 3000110), distal part of penis: A, dorsal view; B, lateral view; C, ventral view; D, apical view. Scale bars = 0.1 mm.