Myrcia densa (De Candolle 1828: 257 ) Sobral (2006: 136)

Santos, Matheus F., Lucas, Eve & Sano, Paulo T., 2018, A taxonomic monograph of Myrcia sect. Sympodiomyrcia (Myrteae, Myrtaceae), Phytotaxa 380 (1), pp. 448-450 : 448-450

7. Myrcia densa (De Candolle 1828: 257) Sobral (2006: 136) ( Figures 2B, 2F , 3E , 5E , 7A , 17 and 21 to 25)

≡ Calyptranthes densa DC. Aulomyrcia densa (DC.) O. Berg (1855 –1856: 37). Type:— BRAZIL. Minas Gerais, 1817 (fr.), Martius 144 (lectotype M [barcode M-0136871]! designated by Santos et al. (2016b), isolectotypes G-DC!, M [barcode M-0136872]!)

= Aulomyrcia supraaxillaris O. Berg (1857 –1859: 69). Myrcia jacobinensis Mattos (1966: 60) . Type:— BRAZIL. Bahia: Jacobina, 1841 (fl.), Blanchet 3368 (lectotype W! designated by Santos et al. (2016b), isolectotypes BM [two sheets]!, C!, F [two sheets]!, G!, HAL [image!], LE!, MICH!, NY [four sheets]!, P [three sheets]!)

Shrub to tree 0.5–6.0 m high. Epidermal peeling sometimes present on immature parts; trichomes brown, light brown to white (rarely ferruginous), 0.1–0.6 mm long. Twig when immature brownish (when dry), flattened, keeled, pubescent, puberulent to glabrous; mature twig greyish (when dry), cylindrical, cortex slightly cracked, glabrescent to glabrous; branching monopodial (rarely sympodial), 2–3 branches per node (rarely more), epidermal protrusion absent at the nodes (present only when branching is sympodial), internode 0.5–4.5 cm long; cataphyll scale-like to foliaceous, 2–5 × 1 mm, usually present only at the basal internode of a new branch, early deciduous, free, ovate, externally puberulent to glabrous, internally glabrous; terminal node with central bud developed and lateral ones undeveloped or central bud undeveloped and lateral ones developed, pubescent, puberulent or with scattered trichomes. Leaf discolorous, chartaceous to coriaceous, blade 0.5–4.5 × 0.2–2.5 cm, narrowly elliptic to circular, ovate or obovate to very widely obovate, apex acuminate to rounded, base attenuate, cuneate to rounded or truncate, margin plane, secondary veins 1–3 mm apart, held at an angle of 35–70° relative to the midvein, marginal vein 0.5–1.0 mm from the margin, tertiary veins inconspicuous; adaxial surface pubescent, puberulent or with scattered trichomes to glabrous when immature, glabrescent to glabrous at maturity, midvein sulcate to flat in the first half and flat in the second half, secondary veins inconspicuous (rarely raised), pellucid dots conspicuous to inconspicuous, less than 5 to more than 15 per mm 2; abaxial surface pubescent, puberulent or with scattered trichomes to glabrous when immature, puberulent or glabrescent to glabrous at maturity, midvein raised, secondary veins inconspicuous or raised, pellucid dots conspicuous to inconspicuous, less than 5 to more than 15 per mm 2; petiole 1–5 × 1–2 mm, canaliculate to semicylindrical, pubescent, puberulent or with scattered trichomes to glabrous when immature, glabrescent to glabrous at maturity. Inflorescence 1–7 × 0.5–8.0 cm, pyramidal or corymbiform (rarely umbelliform), axillar at the terminal or subterminal nodes, terminal dichasia usually with three flowers, 9–67 flowers, rachis pubescent, puberulent or with scattered trichomes to glabrous, 1–2 branching at the base (rarely more than two branching, sometimes with a central vegetative branch), first internode of central rachis 1.0– 1.5 mm wide, semicylindrical to flattened, distal internodes flattened, opposite branching (rarely subopposite), 2–3 (rarely 4–5) branching per node, epidermal protrusion present at the nodes (usually absent in the apical branches). Bract 0.8–5.4 × 0.2–2.8 mm, deciduous, lanceolate, ovate or elliptic, concave or plane, apex acuminate, acute or rounded, base truncate or short petiolate, adaxial and abaxial surfaces pubescent, puberulent or with scattered trichomes to glabrous. Pedicel 0–1.6 mm long, cylindrical, pubescent, puberulent or with scattered trichomes. Bracteole 0.6–2.8 × 0.2–0.6 mm, deciduous, lanceolate or ovate, concave, apex acuminate or acute, base truncate, adaxial surface puberulent or with scattered trichomes to glabrous, abaxial surface puberulent or with scattered trichomes. Floral bud 2–3 × 1–2 mm, turbinate. Hypanthium 0.6–1.0 mm extending above the summit of the ovary, not tearing at anthesis, externally pubescent, puberulent or with scattered trichomes, glabrescent towards the apex, pellucid dots conspicuous (rarely inconspicuous), internally glabrous; calyx 4–5–merous, lobes 0.4–1.6 × 0.4–1.6 mm, distinct from the hypanthium, deciduous, depressed ovate to ovate, concave, apex rounded, base truncate, externally and internally puberulent or with scattered trichomes to glabrous; corolla 4–5– merous, petals light brown to white, 0.8–1.8 × 0.6–2.0 mm, depressed ovate to widely ovate, concave, apex rounded, base truncate, externally and internally puberulent or with scattered trichomes to glabrous; staminal ring 0.1–0.2 mm wide, glabrous, stamens 24–55, filament 1.6–4.0 mm long, white, glabrous, anther 0.24–0.40 × 0.24–0.32 mm, square, oblong or transversely oblong; ovary 0.6–1.0 × 0.8–1.2 mm, 2–locular, each locule with two ovules (rarely 3), style 3.6–5.0 mm long, glabrous, stigma punctiform, papillose. Fruit green when immature, reddish to vinaceous at maturity, 4–6 × 4–6 mm, depressed globose or globose, base rounded, glabrescent to glabrous, remnants of calyx lobes present or not; seeds 1–3.

Distribution and Habitat:— Myrcia densa is mainly distributed in the Espinhaço Range from the Diamantina Plateau to the Chapada Diamantina (up to 2000 m of elevation). It is also found at other high elevation areas, including the municipality of Monte Santo (Bahia state) and the Vale do Catimbau (Pernambuco state), rarely in lowland areas, such as the municipalities of Alagoinhas and Salvador (Bahia state), Igarassu (Pernambuco state), and Alhandra and Santa Rita (Paraíba state) ( Figure 17 ).

The species inhabits different vegetation types in campo rupestre areas as carrasco, cerrado rupestre, grassy fields on sandy soil, riparian forests and rock outcrops. It is occasionally recorded in caatinga ( Filadélfia–Bahia state, and Buíque–Pernambuco state), grassy fields ( Igarassu ), sandy banks in coastal areas ( Salvador – Bahia state) and semideciduous forest ( Alagoinhas ). The distribution of Myrcia densa includes the Atlantic Forest , Caatinga and Cerrado domains .

Phenology:— Myrcia densa was collected in flower from September to March, with flowering peak in November- December and February. Fruits were found from August to September and from December to June (mature fruits in December, February and April).

Conservation Status:— The Area of Occupancy of Myrcia densa is 140 km 2, but this is probably underestimate because of the extremely wide distribution in campo rupestre. Myrcia densa has been recorded in protected areas and sometimes population density is very high. Myrcia densa is here classified as Least Concern (LC; IUCN 2001).

Discussion:— Myrcia densa morphology varies widely but it is well characterized by monopodial vegetative branching, cataphyll scars usually present only at the basal internodes of a new branch, keeled immature twigs ( Figure 3E ), inconspicuous venation and a corymbiform inflorescence with only 1–2 branches at the base ( Figures 5E , 7A ). The species is similar to Myrcia ascendens and Myrcia mucugensis (see comments in these species about their differences).

A peculiar morphological variation of Myrcia densa is found in the region of the Morro do Pai Inácio (Chapada Diamantina – Bahia state): low shrubs with small leaves, very short internodes and immature strongly flattened twigs. Specimens with wider leaves, thick branches and inflorescence rachis are found in the region of Jacobina (Bahia state), the type locality of Myrcia jacobinensis ( Santos et al. 2016b) .

Available illustrations and images:— None found.

Additional specimens examined:— BRAZIL. Bahia: Serra do Sincorá, 1300 m, 16 ° 35’0”S, 41 ° 27’0”W, 22 March 1980 (fr) , R.M.Harley 20734 (CEPEC!, K!, RB!, UB!). Mun.Abaíra, 1800 m, 13 ° 15’0”S, 41 ° 55’0”W, 5 February 1992 (fr) , B. Stannard H51183 (CEPEC!, HUEFS!, K!, SPF!, UB!); ibidem, 1500–1700 m, 13 ° 14’0”S, 41 ° 54’0”W, 26 March 1992 (fr) , E.M. Nic Lughadha H53347 (K!, SPF!); ibidem, 1220 m, 13 ° 17’0”S, 41 ° 52’0”W, 27 December 1988 (fl) , R.M. Harley 27836 (CEPEC!, K!, SPF!, UB!); ibidem, 1350–1650 m, 13 ° 18’0”S, 41 ° 52’0”W, 21 December 1991 (fl) , R.M. Harley 50247 (CEPEC!, HUEFS!, K!, MBM!, SPF!, UB!); ibidem, 1350–1650 m, 13 ° 18’0”S, 41 ° 52’0”W, 21 December 1991 (fl) , R.M. Harley 50248 (K!, SPF!, UB!); ibidem, 1350 m, 13 ° 18’0”S, 41 ° 52’0”W, 26 October 1992 (fl) , W. Ganev 1369 (CEPEC!, ESA!, HUEFS!, K!, SPF!); ibidem, 1250 m, 13 ° 18’0”S, 41 ° 52’0”W, 16 December 1992 (fr) , W. Ganev 1666 (CEPEC!, HUEFS!, K!, SPF!); ibidem, 1950–2033 m, 13 ° 18’0”S, 41 ° 54’0”W, 28 September 1993 (fl) , W. Ganev 2268 (HUEFS!, K!, SPF!, UB!); ibidem, 1950–2033 m, 13 ° 18’0”S, 41 ° 54’0”W, 28 September 1993 (fr) , W. Ganev 2269 (ESA!, HUEFS!, K!, SPF!, UB!); ibidem, 1950–2033 m, 13 ° 18’0”S, 41 ° 54’0”W, 28 September 1993 (fl) , W. Ganev 2276 (HUEFS!, K!, SPF!, UB!); Serra dos Frios, 1700 –1800 m, 13 ° 20’0”S, 41 ° 53’0”W, 12 November 1993 (fl) , W.Ganev 2477 (HUEFS!, K!, SPF!, UB!). Mun.Abaíra–Catolés, Serra do Barbado , 1750–2035 m, 13 ° 17’0”S ,

41 ° 54’0”W, 26 February 1994 (fr), P.T.Sano CFCR14626 (ESA!, K!, SPF!, UB!). Mun. Água Quente , Pico das Almas, 1450 m , 13 ° 31’0”S, 42 ° 0’0”W, 16 December 1988 (fl), R.M.Harley 27510 (CEPEC!, K!, SPF!, UB!). Mun.Alagoinhas , 4 October 2004 (fl), N.G.Jesus 2119 ( HUEFS!). Mun. Andaraí, 710 m , 12 ° 53’4,1”S, 41 ° 18’25,7”W, 11 February 2012 (fr), M.F.Santos 821 (ALCB!, K!, RB!, SPF!); Chapada Diamantina , 12 ° 48’13”S, 41 ° 20’25”W, 7 December 2007 (fl), M.L.Guedes 14190 (ALCB!, MBM!). Mun. Carnaíbas , April 1994 (fr), M.Sobral 7591 (BHCB!, HUEFS!). Mun. Filadélfia, 350 m , 10 ° 46’20”S, 40 ° 8’55”W, 27 February 2000 (fr), A.M.Giulietti 1876 (BHCB!, HUEFS!). Mun. Ibicoara , Chapada Diamantina , 1235 m , 13 ° 25’0”S, 41 ° 31’0”W, 5 April 2000 (fr), M.A.Nóbrega 12 ( SPF!). Mun. Itaberaba , ARIE Serra do Orobó, 960 m , 12 ° 24’27”S, 40 ° 32’5”W, 23 September 2005 (fr), D.Cardoso 841 (BHCB!, HUEFS!); Serra do Orobó , 13 November 1983 (fl), G.C.Pereira Pinto 386/83 (BHCB!, HUEFS!, IBGE!, RB!). Mun. Jacobina, 960 m , 11 ° 9’15”S, 40 ° 48’33”W, 31 March 1993 (fl), M.L. Guedes PCD2717 (ALCB!, HUEFS!, K!, SPF!); ibidem, 680 m , 10 ° 54’2”S, 40 ° 21’55”W, 4 April 1996 (fr), M.L. Guedes PCD2807 (ALCB!, CEPEC!, HUEFS!, K!, SPF!); ibidem, 822–939 m , 11 ° 14’0”S, 40 ° 30’0”W, 31 December 2004 (fl), R.Mello-Silva 2840 ( SPF!); Serra do Tombador , 3 April 1986 (fr), A.M.V.Carvalho 2398 (CEPEC!, ICN!, K!, SP!); ibidem, 750 m , 20 February 1993 (fr), A.M.V.Carvalho 4137 (CEPEC!, HUEFS!, MBM!, RB!, SP!, SPF!). Mun. Lençóis , 12 ° 26’50”S, 41 ° 25’14”W, 21 February 1995 (fr), E. Melo PCD1637 (ALCB!, CEPEC!, HUEFS!, K!, RB!, SPF!); ibidem, 1400 m , 19 December 1984 (fr), R.Mello-Silva CFCR7237 (K!, SPF!). Mun. Monte Santo , 13 February 2011 (fr), G.Costa 480 ( HUEFS!). Mun. Morro do Chapéu, 1300 m , 11 ° 35’27”S, 41 ° 12’29”W, 13 March 1996 (fr), A.A. Conceição PCD2347 (ALCB!, HUEFS!, K!, SPF!); ibidem, 1150 m , 16 February 1971 (fr), H.S. Irwin 32269 (F!, MICH!, MO!, NY!, UB!, US!); ibidem, 1270 m , 11 ° 35’28”S, 41 ° 12’24”W, 7 February 1999 (fl), E.B. Miranda 75 (BHCB!, HUEFS!); ibidem, 1055– 1075 m , 11 ° 34’12”S, 41 ° 9’53”W, 17 November 1999 (fl), E. Melo 3135 (BHCB!, HUEFS!); ibidem, 1073 m , 11 ° 26’24”S, 41 ° 11’39”W, 4 May 2007 (fr), F. França 5662 (BHCB!, HUEFS!); ibidem, 1071 m , 11 ° 26’24”S, 41 ° 11’39”W, 20 May 2007 (fr), J.F.B. Pastore 2095 (BHCB!, HUEFS!); ibidem, 1150 m , 22 February 1993 (fr), J.G. Jardim 61 (CEPEC!, RB!, SP!, UB!); ibidem, 30 August 1990 (fr), J.L. Hage 2331 (CEPEC!, HUEFS!, MBM!, RB!, SP!); ibidem, 1235 m , 11 ° 5’27”S, 41 ° 12’34”W, 28 January 2005 (fr), J. Paula-Souza 4805 (BHCB!, ESA!, K!); ibidem, 14 March 1995 (fr), L.P. Queiroz 4275 (BHCB!, CEPEC!, HUEFS!); ibidem, 883 m , 11 ° 37’30,7”S, 40 ° 59’59,4”W, 9 February 2012 (fl), M.F. Santos 811 (ALCB!, K!, NY!, SPF!); ibidem, 883 m , 11 ° 37’30,7”S, 40 ° 59’59,4”W, 9 February 2012 (fl), M.F. Santos 813 (ALCB!, K!, SPF!); ibidem, 1277 m , 11 ° 35’26,2”S, 41 ° 12’25,5”W, 9 February 2012 (fl, fr), M.F. Santos 814 (K!, SPF!); ibidem, 1277 m , 11 ° 35’26,2”S, 41 ° 12’25,5”W, 9 February 2012 (fl), M.F.Santos 815 (ALCB!, K!, RB!, SPF!); Serra do Tombador , 1150 m , 16 February 1971 (fl), H.S. Irwin 32270 (MICH!, MO!, NY!, US!). Mun. Mucugê, Parque Nacional da Chapada Diamantina , 1388 m , 12 ° 45’27,6”S, 41 ° 30’29,4”W, 12 February 2012 (fl, fr), M.F. Santos 827 (ALCB!, K!, RB!, SPF!). Mun. Palmeiras , 18 January 1997 (fr), A.A. Conceição 294 ( SPF!); 19 February 1997 (fr), A.A. Conceição 412 ( SPF!); 25 January 2000 (fl, fr), A.A. Conceição 730 (HUEFS!, SPF!); ibidem, 25 January 2000 (fr), A.A. Conceição 733 (HUEFS!, SPF!); ibidem, 1140 m , 12 ° 27’30”S, 41 ° 28’23”W, 10 February 2012 (fl, fr), M.F. Santos 820 (ALCB!, K!, RB!, SPF!); ibidem, 1100 m , 12 ° 27’22”S, 41 ° 28’34”W, 27 December 1994 (fl), M.L. Guedes PCD1416 (HUEFS!, K!, RB!, SPF!); ibidem, 28 February 1997 (fr), F.França PCD5911 (K!, US!). Mun. Piatã , 29 November 1999 (fl), A.A. Conceição 631 ( SPF!); ibidem, 11 January 2000 (fr), A.A. Conceição 693 ( SPF!); ibidem, 14 February 1987 (fr), R.M. Harley 24210 (K!, SPF!); ibidem, 1400 m , 13 ° 5’0”S, 41 ° 53’0”W, 5 December 1992 (fl), W. Ganev 1617 (CEPEC!, HUEFS!, K!, SPF!, UB!); ibidem , 13 ° 9’0”S, 41 ° 45’0”W, 20 December 1992 (fr), W.Ganev 1713 (CEPEC!, HUEFS!, K!, SPF!, UB!); Serra do Santana , 1229 m , 13 ° 9’4,5”S, 41 ° 45’52”W, 14 February 2012 (fl, fr), M.F.Santos 835 (ALCB!, K!, RB!, SPF!). Mun. Pindobaçu , Serra da Paciência, 1000 m , 10 ° 56’36”S, 40 ° 24’28”W, 10 April 2001 (fr), N.G. Jesus 1376 ( ALCB!). Mun. Rio de Contas , 13 December 1984 (fl), A.M. Giulietti CFCR6755 (K!, SPF!); ibidem, 9 September 1981 (fr), J.R. Pirani CFCR2176 ( SPF!); ibidem, 1300 m , 13 ° 28’0”S, 41 ° 51’0”W, 8 November 1988 (fl), R.M.Harley 26039 (CEPEC!, ESA!, K!, NY!, RB!, SP!, SPF!, UB!); Pico das Almas, 1450–1500 m , 13 ° 31’0”S, 41 ° 58’0”W, 29 November 1988 (fl), R.M.Harley 26668 (BHCB!, CEPEC!, K!, MBM!, NY!, SP!, SPF!, UB!). Mun. Ruy Barbosa , 12 ° 25’20”S, 40 ° 33’33”W, 14 June 1978 (fr), A.P.Araújo 5 ( RB!); Serra do Orobó , 640–807 m , 12 ° 18’35”S, 40 ° 29’3”W, 18 December 2004 (fl), L.P.Queiroz 9879 (BHCB!, HUEFS!). Mun. Salvador, 18 December 1986 (fl), L.B. Silva 15 ( BHCB!); Abaeté, 17 February 2006 (fl), E.P.Queiroz 1442 ( BHCB!); ibidem, 3 May 1993 (fr), H.O.S.Chagas 2 ( BHCB!); ibidem, 24 January 1965 (fl), R.P.Belém 261 (IAN!, RB!). Mun. Saúde , 645–740 m , 10 ° 56’33”S, 40 ° 24’26”W, 15 May 1999 (fr), F. França 2958 (BHCB!, HUEFS!). Minas Gerais : Mun. Grão-Mogol, 7 January 1986 (fr), C. Kameyama CFCR9010 (NY!, SP!, SPF!, UEC!); ibidem, 21 October 1978 (fl), G. Hatschbach 41600 (BR!, C!, MBM!, SP!); ibidem, 22 October 1978 (fl), G. Hatschbach 41650 (MBM!, SP!); ibidem, 7 January 1986 (fr), I. Cordeiro CFCR9007 (NY!, SP!, SPF!, UEC!); ibidem, 695 m , 16 ° 35’25,6”S, 42 ° 54’55,2”W, 10 February 2011 (fr), M.F. Santos 643 (BHCB!, K!, NY!, RB!, SPF!); ibidem, 1000–1200 m , 11 December 1989 (fr), P.T. Sano CFCR12469 (K!, NY!, SPF!); ibidem, 650 m , 16 ° 36’0”S, 42 ° 55’0”W, 12 December 1989 (fr), P.T. Sano CFCR12564 (K!, NY!, SP!, SPF!); Parque Estadual de Grão-Mogol , 710 m , 16 ° 35’43,5”S, 42 ° 57’18,1”W, 11 February 2011 (fl), M.F.Santos 644 (K!, SPF!). Mun.Itacambira , 1250m , 16 ° 59’6,16”S, 43 ° 20’42”W, 13 November 2001 (fl), A.M.G.A.Tozzi 2001-442 (BHCB!, UEC!). Mun. Jequitinhonha , 1080 m , 16 ° 20’21”S, 41 ° 5’24”W, 7 February 2005 (fr), J. Paula-Souza 5611 (BHCB!, ESA!, K!); Reserva Biológica da Mata Escura , 1134 m , 16 ° 20’18”S, 41 ° 5’23”W, 25 March 2008 (fr), T.E.Almeida 1315 (BHCB!, MBM!). Paraíba: Mun. Alhandra, 15 December 1976 (fr), M.C.R.Tavares 1 ( RB!). Mun. Santa Rita, 8 February 1968 (fl), Andrade-Lima 68-5316 ( RB!). Pernambuco: Mun. Buíque, 14 February 2008 (fl), G.D.Alcântara 44 (HUEFS!, SPF!); ibidem, 1 November 1961 (fl), Andrade-Lima 61-3986 (K!); ibidem, 799 m , 12 January 1996 (fr), K. Andrade 297 ( NY!); ibidem, 700 m , 8 ° 37’S, 37 ° 10’W, 1 October 1994 (fl), M.J.N.Rodal 412 (K!, RB!). Mun. Igarassu , 19 October 1887 (fl), G.A.Ramage s.n. ( BM!). Total: 89 specimens .

Figures

FIGURE 2. Habits of Myrcia sect. Sympodiomyrcia. A. Shrub, Myrcia mutabilis (Santos 591 [SPF]). B. Shrub, Myrcia densa (Santos 820 [SPF]). C. Tree, Myrcia subcordata (Faria 3105 [UB]). D. Shrub, Myrcia rupestris (Santos 654 [SPF]). E. Shrub, Myrcia mucugensis (Santos 823 [SPF]). F. Treelet, Myrcia densa (Santos 813 [SPF]). G. Subshrub, Myrcia nitida (Santos 831 [SPF]). H. Shrub, Myrcia ascendens (Santos 829 [SPF]). Image credit: C. Jair E.Q.Faria.

FIGURE 3. A. Detail of the leaf of Myrcia summa showing foveolate dots (Prance 29040 [RB]). B. Wax peeling (white) on the leaf of Myrcia rupestris (Santos 641 [SPF]). C. Wax peeling (white) on the leaf of Myrcia costeira (Santos 851 [SPF]). D. Keeled twig of Myrcia costeira (Santos 851 [SPF]). E. Keeled twig of Myrcia densa (Santos 643 [SPF]). F. Leafy cataphyll of Myrcia subcordata covering immature leaves (Rochelle 55 [SPF]). G. Leafy cataphyll (indicated) of Myrcia costeira (Santos 851 [SPF]). H. Scale-like cataphyll (indicated) of Myrcia subavenia (Santos 715 [SPF]). Scales:A=4 mm; B–H=1 cm.

FIGURE 5. Inflorescence of Myrcia sect. Sympodiomyrcia. A. Myrcia bicolor, sympodial branching at the base of the inflorescence, with terminal dichasia (Ávila 388 [SPF]). B. Myrcia tenuifolia, multiple branches at the base, including also vegetative branches (Santos 1223 [ICN]). C. Myrcia rupestris, inflorescence with one branch per axillary node, terminal dichasia present (Santos 640 [SPF]). D. Pyramidal inflorescence of Myrcia plusiantha (Barbosa 931 [SPF]). E. Corymbiform inflorescence of Myrcia densa (Santos 813 [SPF]). F. Umbelliform inflorescence of Myrcia mucugensis (Kral 75641 [SP]). Scales: C=1 cm; F=5 mm.

FIGURE 7. Fruits of Myrcia sect. Sympodiomyrcia. A. Myrcia densa (Santos 643 [SPF]). B. Myrcia mucugensis (Santos 823 [SPF]). C. Myrcia ascendens (Santos 829 [SPF]). D. Myrcia mutabilis (Faria 3106 [UB]). Image credit: D. Jair E.Q.Faria.

FIGURE 17. Distribution of Myrcia bicolor (circles) and Myrcia densa (squares). Continuous lines represent South American country and Brazilian state boundaries. The grey gradient represents elevation.

FIGURE 21. Images of representative specimens of Myrcia sect. Sympodiomyrcia species: Myrcia densa (Santos 643 [SPF]).