Harmonia Mulsant, 1846

Biranvand, Amir, Nedvěd, Oldřich, Tomaszewska, Wioletta, Al Ansi, Amin N., Fekrat, Lida, Haghghadam, Zahra Mojib, Khormizi, Mehdi Zare, Noorinahad, Sara, Şenal, Derya, Shakarami, Jahanshir & Haelewaters, Danny, 2019, The genus Harmonia (Coleoptera, Coccinellidae) in the Middle East region, Acta Entomologica Musei Nationalis Pragae 59 (1), pp. 163-170 : 164

Harmonia Mulsant, 1846

= Leis Mulsant, 1850 ; Ballia Mulsant, 1853 ; Callineda Crotch, 1871 ; Stictoleis Crotch, 1874 ; Ptychanatis Crotch, 1874 ; Rhopaloneda Timberlake, 1943 (sensu ŚLIPIŃSKI 2007).

Diagnosis. Length 4–8 mm. Elytra moderately to strongly convex, glabrous; elytral color pattern variable. Clypeus anteriorly straight between lateral projections. Antenna composed of 11 antennomeres, slightly shorter than the width of head capsule, with club moderately compact, composed of three antennomeres ( Fig. 9 ). Terminal maxillary palpomere securiform ( Fig. 11 ). Pronotal disc evenly convex, transverse with narrowly upturned and/or slightly thickenedexternal borders. Prothoracichypomeronwithout fovea near anterior angles; prosternal process without or withdistinctlateral carinae extendinganteriorly ( Figs 6–7 ). Anteriormarginof mesoventriteweaklyfoveate mediallyto straight. Elytral lateral marginsnarrowly raised; epipleuron not foveate. Apices of middle and hindtibiae without spurs ( Fig. 8 ). Abdominalpostcoxallinesnot recurved, incomplete laterally ( Fig. 12 ). Penis stout, consists of basal sclerite and additionalapical piece. Ovipositor withcoxitesclub-handle type; infundibulum present; spermatheca C-shaped.

Figures

Figs 6–13. Morphological characters of Harmonia species: 6 – H. axyridis (Pallas, 1773), prosternum. 7–13 – Harmonia quadripunctata (Pontoppidan, 1763): 7 – prosternum; 8 – hind leg; 9 – antenna; 10 – meso- and metaventrites; 11 – maxillary palp; 12 – abdominal ventrite 1 with postcoxal lines; 13 – mandible.