Latarima, Shackleton & Webb & Lawler & Suter, 2014

Shackleton, M. E., Webb, J. M., Lawler, S. H. & Suter, P. J., 2014, A new genus and species of Calocidae (Trichoptera: Insecta) from south eastern Australia, Memoirs of Museum Victoria 72, pp. 25-30 : 26-29

Latarima Genus nov.

Zoobank LSID. http://zoobank.org/ urn:lsid:zoobank.org:act:

955AF7F6-958F-4485-AC37-80DD885EE461

Type species: L. furcilla (Neboiss) comb. nov. ( Tamasia furcilla

Neboiss in Neboiss 1984a).

Generic Diagnosis. Adult and larval characters can be used to distinguish Latarima from all other Calocidae genera. This genus is the only Calocidae genus known where segment X, in the adult males, is widely separated at the base of the segment. The dorsal surface of the head capsule, in the adult male, does not possess eversible scent organs, as in Caloca and Pliocaloca . The segments of the maxillary palps are expanded, similar to Tamasia , Caenota , and Pycnocentrella . However, unlike Caenota , segments 3 and 4 are not greatly reduced. As with Tamasia , but no other Calocidae genus, there are only 4 maxillary palpal segments.

In the larva, the frontoclypeus widens suddenly towards the anterior margin, unlike those of Caloca , Tamasia , and Calocoides . The posterior and anterior portions of the frontoclypeus are separated by a constriction and the lateral margins of the anterior portion are somewhat rounded. This is dissimilar to Pycnocentrella eruensis where the anterior portion diverges out from the posterior portion in a straight line without any constriction between the two. The head and pronotum is reticulate in texture and does not possess the dense, short, papillate setae present in Tamasia or Pliocaloca . The metanotum does not possess a small, sclerotised ridge on the anterior margin, as in Caenota and Calocoides . The foretrochantin is fused to the propleuron, unlike that of Pliocaloca .

Generic description. Adult male: Dark brown to blackish, white markings on wings. Head: dorsum with single setal wart along midline; small receptacle on posterior margin dorsally extending mid-dorsal setal wart, not membranous, without eversible scent organs; posterior setal warts absent; postocular setal warts long, narrow. Maxillary palps four segmented, segments irregular, bulbous, apical segment reflexed posteriorly; labial palps three segmented; antenna slightly shorter than fore wing; scape long, with highly setose anterior projection. Pronotum: one pair lateral setose warts. Mesoscutellum smooth. Scutellum with one pair, elongate, setal warts. Legs: tibial spines 2:2:4. Wings: Forewing: discoidal cell present; thyridial cell present; fork 1 and 2 sessile; fork 3 petiolate; fork 4 absent; fork 5 sessile or slightly petiolate. Hind wing: venation reduced, with or without posterior fold; discoidal cell present; thyridial cell absent; base of M absent; large vein-free area in basal posterior section if no fold present; Sc and R1 fused; cell formed by Sc and C relatively large; fork 1 sessile, no other forks present. Genitalia: Segment X divided into two elongated, widely separated segments.

Larva: Head: reticulate in texture; frontoclypeus widens suddenly anteriorly; antennae close to eye. Pronotum: with weak lateral carina; foretrochantin fused to propleuron. Metanotum: single sclerite in anterior half. Abdomen: segment 1 lateral hump with large spiny patch, without sclerites; gills absent.

Etymology. From the Latin lata meaning wide and rima meaning gap, and pertaining to the widely incised segment X and is feminine in gender.

Material examined. Species included: Latarima explicatala sp nov. and Latarima furcilla ( Neboiss, 1984a) comb. nov.

Comments. Adults of Latarima furcilla have only been collected from a single stream on the road to Mt Buller, Victoria. Preliminary investigations suggest that a larva from the same site is a likely candidate for being associated with the adults of this species. However, this association has not yet been confirmed. The range of L. explicatala is much wider than that of L. furcilla . Latarima explicatala has been collected from the Yarra Ranges, Mount Baw Baw, Taggerty, Mount Buller, and at the Victorian-New South Wales border near Mt. Kosciuszko. These sites are all well forested and associated with mountains. Interestingly, one site where this L. explicatala has been collected from is less than 1km from the site at which L. furcilla is found, on Mt Buller.