Mortoniella (Mortoniella) enchrysa Flint, 1991

Blahnik, Roger J. & Holzenthal, Ralph W., 2017, Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *, Insecta Mundi 2017 (602), pp. 1-251 : 28-29

Mortoniella (Mortoniella) enchrysa Flint, 1991

Fig. 13

Mortoniella enchrysa Flint 1991: 24 ; Sykora 1999: 386 [member of enchrysa subgroup]; Blahnik and Holzenthal 2008: 70 [member of bilineata group]; Blahnik and Holzenthal 2011: 63 [member of enchrysa subgroup].

Mortoniella enchrysa Flint is most similar overall to M. adamsae , n. sp., especially in the shape of the inferior appendages, but those of M. enchrysa have the apical processes shorter, more strongly arched, and densely covered with microsetae. Both species have spines or spine-like processes on the paramere appendages, but M. enchrysa has only one pair of appendages. An additional similarity between M. enchrysa and M. adamsae is in the structure of the ventral lobes of the phallicata, which have the basal part forming retrorse lobes. The species differ in that the apical notch of tergum X is shallower in M. enchrysa . Mortoniella enchrysa also differs in that the posterior margin of segment IX is very sharply, nearly rectilinearly, angular. The latter character state is similarly developed in M. silacea , n. sp., but there is otherwise no close resemblance between the two species.

Adult —Length of forewing: male 6.0 mm; female 6.8. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0:4:4. Dorsal side of forewings, head, basal segments of antennae, and legs golden-orange; wing membrane of fore- and hind wings distinctly infuscated, ventral side of forewings (and apicomarginal setae), hind wings, apices of antennae, and palps dark brown. Tibial spurs brownish-black, strongly contrasting with legs. Wing bars absent.

Male genitalia —Ventral process of segment VI posteriorly projecting, short, narrow basally, length about 1½ to 2 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX, membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and produced in ventral half, posterolateral margin with nearly rectilinearly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X elongate, lateral margins rounded, ventrolaterally with subangular lateral lobes, each with prominent apical seta; apex of tergum distinctly sclerotized, emarginate, with ventrolateral margins incurved and approaching each other mesally, but separated by distinct gap, apicodorsally with lightly sclerotized connection near apex (mesal notch shallow, but distinct); tergum ventromesally with paired, sclerotized, rounded (short paddle-like), ventromesal lobes in basal half, each with short setae. Inferior appendages with short upright dorsolateral lobes and short paired ventromesal lobes, ventromesal lobes strongly curved, each with distinct preapical spine on ventral margin; basodorsal lobes and base of ventromesal lobes with numerous microsetae. Mesal pockets of inferior appendage with moderately elongate, posteriorly-directed, spine-like, apicoventral projections. Paramere appendage elongate, narrow, extending about same length as dorsal phallic spine, distinctly dorsally curved and slightly widened near apex, apex acute; ventral margin of appendage with linear row of sensilla; fused basal segments of parameres articulating near base of dorsal phallic spine. Phallobase with rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin arched in middle, sinuously and nearly rectilinearly upturned in apical ¼, apex of spine rounded; base of spine narrow and stalk-like, abruptly and strongly widened on ventral margin in basal ½, forming subangular ventral projection, spine narrowing apically from projection; spine, as viewed dorsally, somewhat widened in middle, apex rounded. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine; phallicata ventrally with 2 pairs of diverging, sclerotized lobes, basal ones retrorsely oriented, rounded laterally, posterior pair diverging from posterior margin of basal lobes, apices bluntly rounded, posteriorly projecting; ventromesal margin of phallicata extending beyond paired lobes, very lightly sclerotized, extending about same length as paramere appendages. Endophallic membrane simple in structure, with very small membranous preapical lateral lobes; phallotremal spines absent.

Material examined — COLOMBIA: Valle: Municipio El Cerrito, Río Cerrito, 7.1 km E Hacienda “El Paraiso”, 3.64972° N, 76.17278° W, 1950 m, 3.xii.1997, F Muñoz et al.– 1 male, 1 female (pinned) (UMSP).

Distribution — Colombia.

Figures

Figure 13. Mortoniella (Mortoniella) enchrysa Flint, male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral; F—membranous connection between segments VIII and IX.