Dichotomius (Selenocopris) danieli Kohlmann and Solís, 1997

Valois, Marcely C., Silva, Fernando A. B. & Vaz-de-Mello, Fernando Z., 2022, A taxonomic revision of the globulus species group of Dichotomius Hope, 1838 (Coleoptera: Scarabaeidae: Scarabaeinae), Journal of Natural History 56 (1 - 4), pp. 119-147 : 131-132

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https://doi.org/ 10.1080/00222933.2022.2046887



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Dichotomius (Selenocopris) danieli Kohlmann and Solís, 1997


Dichotomius (Selenocopris) danieli Kohlmann and Solís, 1997 View in CoL

( Figures 2 View Figure 2 (a–f), 6))

Dichotomius danieli Kohlmann and Solís 1997: 345 View in CoL , 362; Krajcik 2006: 53; 2012: 91; Solís and Kohlmann 2012: 6; Vaz-de-Mello and Nunes 2016: 299.


Dichotomius danieli , D. rodrigoi and D. favi are the only known species of the globulus group recorded in Costa Rica. As pre-apical teeth are absent on the dorsal surface of paramera, D. danieli is suggested to be more closely related to D. favi than to D. rodrigoi . The following characters can be used to distinguish D. danieli from D. favi and other species in the globulus group: surface of head covered with dense ocellate punctures; pronotal punctures equally spaced from each other; anteromedian region of pronotum with a feeble central impression; elytral striae wide, crenulate, punctures almost as wide as striae; elytral interstriae opaque with indistinct smooth dots; paramera rectangular, apex rounded in dorsal view ( Figure 2 View Figure 2 (d)).

Redescription (male)

Body length: 9.5 mm. Dorsum black, barely shiny ( Figure 2 View Figure 2 (a)). Head: clypeal edge bidentate, teeth acute, separated by a deep V-shaped emargination. Clypeogenal junction rounded. Surface with different-sized ocellate punctures; punctures forming transverse rows at anterior clypeal region. Clypeogenal suture present. Cephalic process produced into a small central tubercle ( Figure 2 View Figure 2 (a)). Antennal club with light brown tumescence. Thorax: pronotum approximately twice wider than long. Pronotal disc convex, anteromedian region with a shallow central excavation; surface with ocellate punctures as dense as those on anterior angles ( Figure 2 View Figure 2 (a)). Surface of hypomera with sparse, coarse setiferous punctures; setae short, and usually absent in some punctures. Prosternum filled with ocellate punctures. Mesoventrite filled with ocellate punctures usually bearing short, orange setae. Metaventrite surface with ocellate punctures, except on disc; punctures on anterior region usually bearing short setae. Meso-metaventral suture distinct. Mid-longitudinal sulcus feebly marked posteriorly. Lateral lobes of metaventrite with ocellate punctures larger than those on anterior region. Elytra: slightly convex in lateral view. Humeral callus distinct. Striae thin, with well-defined punctures spaced by their diameter. Interstriae I–VII slight convex, barely shiny, with shagreened microsculpture; medial region with distinct smooth dots. Legs: protibiae with three distinct teeth externally. Protibial spur curved at apical fifth. Metatibial spur deeply emarginate apically. Ventral surface of the profemora covered with coarse setiferous punctures along its entire extension, setae short. Ventral surface of the mesofemora with ocellate punctures at apical half. Ventral surface of the metafemora with ocellate punctures concentrated on apical half; basal half with sparse, scattered punctures. Meso and metafemora bearing a thin longitudinal groove along the entire extension of posterior margin, wider on apical half. Abdomen: ventrites covered with ocellate punctures laterally, punctures medially disposed in a single row along anterior edge. Pygidium twice wider than long, disc strongly convex apically. Surface of pygidium with ocellate punctures concentrated along anterior half, and spaced by half their diameter. Apex of the pygidium margined. Tegmen: in dorsal view, paramera rectangular, one-third longer than its width medially, apex truncate; inner edges of paramera curved outward along its whole length ( Figure 2 View Figure 2 (d)). Dorsal surface of paramera smooth ( Figure 2 View Figure 2 (d)). Apical half noticeably narrower than base in lateral view ( Figure 2 View Figure 2 (f)).Ventral sclerotised membrane between paramera thinner and curved at basal half; rounded laterally ( Figure 2 View Figure 2 (e)). Female: differs from male by clypeal teeth acute and separated by a deep V-shaped emargination, and sixth abdominal ventrite 3 times longer than fifth.

Type material examined

Holotype of Dichotomius danieli ♂ [ MNCR]: ‘Est. Pitilla, 9 km S. Santa Cecilia, Prov./Guana, COSTA RICA, 700 m. Ago 1994 /C. Moranga, L N 330200_380200 # 3198’ // ‘ Dichotomius danieli /Kohlmann and Solís’/‘ COSTA RICA INBIO/[barcode] CRI002/029748’ // ‘HOLOTYPO/ Dichotomius danieli /Kohlmann y Solís’. Examined via photographs ( Figure 2 View Figure 2 (c)).


Museo Nacional de Costa Rica














Dichotomius (Selenocopris) danieli Kohlmann and Solís, 1997

Valois, Marcely C., Silva, Fernando A. B. & Vaz-de-Mello, Fernando Z. 2022

Dichotomius danieli Kohlmann and Solís 1997: 345

Vaz-de-Mello FZ & Nunes RV 2016: 299
Krajcik M 2012: 91
Solis A & Kohlmann B 2012: 6
Krajcik M 2006: 53
Kohlmann B & Solis A 1997: 345
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