Procladius Skuse, 1889

Chamutiová, Tímea, Hamerlík, Ladislav & Bitušík, Peter, 2020, Subfossil chironomids (Diptera, Chironomidae) of lakes in the Tatra Mountains an illustrated guide, Zootaxa 4819 (2), pp. 216-264 : 227

Procladius Skuse ( Fig. 26 )

Large, oval head capsule whitish to yellowish in colour. Mandible slender, uniformly curved with dark black tip of the apical tooth; mola distally expanded to a large, broad tooth with bluntly rounded apex ( Fig. 26 ). Dorsomental teeth (6–8) present. Ligula with 5 dark teeth, their apex is blackish brown. Paraligula multibranched with more or less prominent main spine ( Fig. 26A ). Shape of paraligula is diagnostic to differ the subgenera Holotanypus and Psilotanypus ( Cranston & Epler 2013) . Pecten hypopharyngis with 10–15 teeth and some smaller teeth in a supplementary row.

Remarks: Procladius remains differ from all other Tanypodinae with dorsomental teeth, in blackish-brown coloration of distal part of ligula and apex of mandibula, and in large, blunt mandibular mola expansion.

Detailed survey based on pupal exuviae study revealed presence of two species in the Tatra Mts.: P. (Holotanypus) choreus (Meigen, 1804) and P. (Holotanypus) tatrensis Gowin, 1944 . While P. tatrensis is widespread with exception of the coldest lakes, P. choreus was repeatedly found in one, strongly acidified lake ( Bitušík et al. 2006a).

Subfossil remains were common in most of studied lakes. All of them belong to subgenus Holotanypus .

Figures

FIGURES 24–27. Ablabesmyia longistyla-type: 24—head capsule (arrows indicate thorn-like projection and segmented maxillary palp), A—maxillary palp, B—pecten hypopharyngis. Macropelopia nebulosa-type: 25—head capsule, A—detail of ligula. Procladius: 26—head capsule (arrow indicates broad tooth on the mandible), A—detail of ligula (arrow indicates paraligula). Zavrelimyia: 27—head capsule (arrow indicates thorn-like projection).