Henderson, Aaron C., Reeve, Alan J., Jabado, Rima W. & Naylor, Gavin J. P., 2016, Taxonomic assessment of sharks, rays and guitarfishes (Chondrichthyes: Elasmobranchii) from south-eastern Arabia, using the NADH dehydrogenase subunit 2 (NADH 2) gene, Zoological Journal of the Linnean Society 176 (2), pp. 399-442: 431

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http://doi.org/ 10.1111/zoj.12309

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Three Sphyrna species   were examined during the present study, namely S. lewini   , S. zygaena   and S. mokarran   , and each formed a separate cluster with a mean between-group distance of 15.10 ± 0.12%. Naylor et al. (2012) reported two main clusters within S. lewini   , designated S. lewini   1 (Atlantic and Indian Ocean specimens) and S. lewini   2 (Pacific specimens). Specimens of S. lewini   from the present study clustered with S. lewini   1, within which they grouped most closely with other Indo-Pacific specimens. Similarly, Naylor et al. (2012) found two main clusters within S. mokarran   , representing Atlantic ( S. mokarran   1) and Pacific ( S. mokarran   2) specimens. The two S. mokarran   speci- mens from the present study clustered with S. mokarran   1. Lastly, only one cluster was reported for S. zygaena   by Naylor et al. (2012), and the addition of the current specimens did not alter this arrangement.