Dichotomius mamillatus (Felsche, 1910)

Cárdenas-Bautista, Johann Stephens, Parada-Alfonso, Jenny Andrea & Carvajal‑Cogollo, Juan E., 2020, Dung beetles (Scarabaeidae, Scarabaeinae) of the Foothills-Andean Forest strip of Villavicencio, Colombia, Check List 16 (4), pp. 821-839 : 830

Dichotomius mamillatus (Felsche, 1910)

Materials examined. COLOMBIA • 1♀, 23.74 mm (mean); Meta, Villavicencio, Vereda El Carmen, “La Loma” Farm ; 04°08′28.18″N, 073°40′01.90″W; 730 m a.s.l.; 18 Oct. 2018; Cárdenas Johann and Parada Jenny leg.; wooded pasture; Crpat 3; UPTC-In-00054 . • 1♂, 23.74 mm (mean); Meta, Villavicencio, Vereda El Carmen, Caño Blanco–Caño Buque ; 04°08′21.85″N, 073°40′12.19″W; 774 m a.s.l.; 17 Oct. 2018; Cárdenas Johann and Parada Jenny leg.; riparian forest; Crbrt 4; UPTC-In-00055 .

Identification. This species can be distinguished from other Dichotomius species by the head with transverse roughness in the clypeus, gena and frons, the presence of a short, thick and blunt horn at the height of the gena with two small horns adjacent to the base of the central horn; pronotum with two small protuberances in the anterior medial region ( Sarmiento-Gárces and Amat-García 2014).

Distribution. The species is distributed in the Colombian and Ecuadorian Amazon, where it is a very wellrepresented species, extending to the foothills up to 1300 m a.s.l. ( Sarmiento-Gárces and Amat-García 2009; Sarmiento-Gárces and Amat-García 2014). In the study area, it was observed between 750–850 m a.s.l. and seen in the habitats of riparian forest, high secondary forest, and wooded grassland, although it was rare species (59 individuals).