Syzeuctus maculipennis ( COSTA , 1883)

Riedel, Matthias, 2021, The genus Syzeuctus FÖRSTER in the Western Palaearctic region and Central Asia (Hymenoptera, Ichneumonidae, Banchinae), with descriptions of 17 new species, Linzer biologische Beiträge 52 (2), pp. 1013-1098 : 1057-1058

Syzeuctus maculipennis ( COSTA, 1883) (Figs 41, 80, 125)

O r i g i n a l: Lissonota maculipennis COSTA, 1883: 101

syn. Lissonota lunigera BRAUNS, 1888: 2, synonymized by YU & HORSTMANN 1997.

P a r a L e c t o t y p e: (♂) " Sierre 15.7.84 Friese " (Berlin), studied. Lectotype not found in Berlin. syn. Lissonota ducalis COSTA, 1890: 16 , synonymized by YU & HORSTMANN 1997 .

syn. Lissonota multipicta KRIECHBAUMER, 1895: 264-266, synonymized by YU & HORSTMANN 1997.

P a r a L e c t o t y p e: (♂) "Sièrre 21-23.6.79. Frey-G.; 830; multipicta m. ♂ " (Bern), studied by Horstmann .

In Munich 1♂ from Sierre labeled as " holotype " by AUBERT in 1973 and 3♂♂ labeled as paralectotypes by HORSTMANN in 2002.

syn. Syzeuctus braunsii SZÉPLIGETI, 1899: 31 , synonymized by YU & HORSTMANN 1997.

H o l o t y p e: (♂) " Budapest Pável ; Holotypus Syzeuctus braunsii SZÉPL. (recte: lunigera) J.F. Aubert 1976" (Budapest), type studied .

S t u d i e d m a t e r i a l: 25♀♀, 35♂♂; Azerbaijan *, Bulgaria, Croatia, France, Greece *, Hungary, Iran *, Kazakhstan *, Switzerland, Turkey *.

D e s c r i p t i o n: ♀: Body length 10.5-12 mm. Antenna with 37-41 flagellomeres; 1 st flagellomere 3.6-4.0× longer than wide and 0.42-0.50× as long as eye; preapical flagellomere 1.1-1.2× longer than wide. Temple moderately and roundly narrowed behind eye, dorsally 0.4× as long as eye. OED 1.5-1.8× and OOD 2.1× ocellar diameter. Frons densely punctate, slightly swollen laterally, with median impression. Face densely punctate, with parallel sides. Clypeus 1.8-2.2× wider than long, with scattered punctures, apical margin rounded. MI 0.6-0.8. Genal carina reaching mandibular base, slightly bent outwards ventrally.

Mesosoma 1.3× longer than high. Frontoventral edge of pronotum moderately curved. Lateral pronotal groove smooth. Epicnemial carina slightly elevated, with ± distinct denticular projection on transversal part. Mesoscutum, mesopleuron including speculum and metapleuron densely punctate. Propodeum with complete apical transversal carina, rarely carina shortly obsolete medially. Hind femur 4.7-5.3× longer than wide. Hind metatarsus 0.7× as long as hind femur. 2 nd hind tarsomere 1.2-1.3× as long as 3 rd one. Hind claw with dense short teeth, slightly curved apically. Areolet stalked; 2 nd recurrent vein in or distal to its middle. Nervulus postfurcal by 1-2× its width. Hairs on basal cell about as long as their intervals. Hind wing with 9-10 distal hamuli.

1 st tergite 1.5-1.9× longer than wide, with scattered or dense coarse punctures. 2 nd tergite 1.0-1.3× longer than wide, densely punctate laterally. 3 rd tergite 1.0-1.3× longer than wide, with dense punctures. Spiracles of 1 st to 3 rd tergites ± protruding laterally. Ovipositor 1.1-1.5× longer than fore wing, 1.3-1.8× longer than metasoma and 2.7-3.4× longer than hind tibia.

Color: Black. Basal spot on mandible, clypeus, inner and outer orbits confluent on vertex, often malar space or complete orbits, sometimes paramedian stripes on face, spot on collare yellow. Mesosoma variable: Dark form (f. lunigera): sometimes anterolateral spot on scutellum, spot on collare, anterolateral spot on mesoscutum and subtegular ridge yellow. Pale form: Mesoscutum anterolaterally and centrally, collare, tegula, subtegular ridge, stripe or spots on mesopleuron, spot on metapleuron, scutellum and postscutellum, and lateral and central spots on propodeum yellow. Tegula yellowish or reddish. Metasoma sometimes black and basal tergites diffusely reddish, usually 1 st- 3 rd tergites red, sometimes 1 st- 4 th tergites with apical yellow bands. Legs except black coxae and trochanters reddish; sometimes coxae and trochanters with yellow spots; sometimes fore and mid legs with yellow frontal stripes; all tibiae narrowly yellowish at base; sometimes hind tibia yellow externally mid and hind tarsi ± brownish or black. Fore wing with (sometimes weak) subapical brown spot; pterostigma brown or black, yellow proximally. ♂: Body length 9-11 mm. Antenna with 36-43 flagellomeres; 1 st flagellomere 3.3-4.0× as long as wide and 0.50-0.55× as long as eye. Temple shortly parallel or slightly rounded behind eye, strongly narrowed apically, dorsally 0.4-0.6× as long as eye. OED 1.6-1.7× and OOD 2.0× ocellar diameter. Clypeus 1.7-2.0× wider than long. MI 0.8-0.9. Genal carina reaching mandibular base.

Lateral pronotal groove smooth ventrally. Epicnemial carina low, with low carina or weak lamella in transversal part. Speculum punctate. Apical transversal carina of propodeum complete. Hind femur 4.3-4.7× longer than wide. Hind metatarsus 0.7-0.75× as long as hind femur. 2 nd hind tarsomere 1.3× longer than 3 rd one. Hind claw with short teeth, moderately bent apically. Areolet stalked; 2 nd recurrent vein slightly proximal or distal to its middle. Nervulus postfurcal by 1-2× its width. Hairs on basal cell about as long as their intervals. Hind wing with 8-9 distal hamuli.

1 st tergite 1.5-1.8× longer than wide, with rather dense punctures. 2 nd tergite 0.9-1.1× as long as wide, with dense punctures. 3 rd tergite 0.9-1.1× as long as wide, with dense punctures. Spiracles of basal tergites not or slightly protruding.

Color: Black and yellow. Face mainly or entirely yellow. Scape and pedicel often yellow ventrally flagellum reddish or blackish. Mesosoma with extended yellow coloration. Basal tergites red or blackish with ± distinct reddish suffusion; 1 st to 2 nd /4 th tergites usually with apical yellow bands. Fore and mid legs largely yellow, sometimes femora and tibiae with red stripes ventrally. Hind coxa black, with yellow spot; hind trochanter usually black, hind trochantellus yellow. Hind femur red or yellow with red dorsal stripe; hind tibia red, yellow basally or externally; mid and hind tarsi ± brown. Subapical spot of fore wing sometimes weak or absent; pterostigma ochreous or blackish, yellow proximally.

Hosts unknown.