Tetraphlebia eleates ( Weymer, 1890 ) (Pseudomaniola)

Matz, Jess & Brower, Andrew V. Z., 2016, The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes, Zootaxa 4125 (1), pp. 1-108 : 70-71

Tetraphlebia eleates ( Weymer, 1890) (Pseudomaniola) n. comb.

( Figs. 12 C, D; 21D–F; 30)

Type location: Tacora, Potosí Province, Bolivia

Holotype: (male) MFN, Berlin, genitalia vial M-9009 Lee D. Miller, 3042 (photo examined) Other combinations:

Faunula eleates Pyrcz (2012)

Faunula leucoglene eleates —Peña & Ugarte (1997, p. 281); Lamas & Viloria (2004, p. 216) Neomaniola eleates —D’Abrera (1988, p. 843)

Distribution. Found in Bolivia throughout Potosí Department from October to December at 4000–5000m ( Fig. 30).

Diagnosis. Most similar to T. leucoglene , but generally lighter in color and with the apical ocellus on the ventral side of the hindwings much reduced. This ocellus is completely contained within M1-M2 and the pupil is a small white point at the center. Males are without androconia visible on the forewing and male genitalia have a narrower uncus, wider gnathos, and a tegumen wider along the dorsoventral side axis than in T. leucoglene . Valvae are deltoid at the distal end in T. eleates and rounded at the distal end in T. leucoglene . Specimens found in Bolivia at high altitudes are more likely to be T. eleates while T. leucoglene is found in Chile much farther south and at lower altitudes.

Redescription. Head: Antennae 6–7mm with dark chocolate scales and a longitudinal stripe of white scales, terminating in a round club. Eyes round and naked, length approximately 1.2 times width. Palps chocolate brown with dark chocolate piliform scales in the males and in the females tan to chocolate with chocolate piliform scales on the dorsal side and dark chocolate to black piliform scales on the ventral side. Terminal segment conical and about one-eighth the length of the second segment. Thorax with iridescent black scales and chocolate to dark chocolate piliform scales in the males and the females with white and iridescent black scales and chocolate piliform scales. Foreleg tarsi with two segments in the males and three segments in the females. Midlegs and hindlegs with four rows of amber spines on the tibia and tarsus.

Forewing ( Fig. 12 C, D): Wingspan 20–22mm. Termen slightly convex to nearly straight and the distal end of the discal cell sinuate with the cubital end straighter than the radial end. Males with no visible androconial patch. Dorsal side chocolate to dark chocolate brown, the males darker. Postmedian band appears in both sexes as a pair of rust orange patches between M3-CuA1 and CuA1-CuA2 with another patch sometimes appearing between M2- M3. Fringe scales are chocolate to dark chocolate. Ventral side similar in color to the dorsal side. Postmedian band rust orange and broken by the veins in some specimens, widest where it surrounds the M1-M2 ocellus, and narrowing toward the tornus. Borders of the postmedian band are dark chocolate to dark coffee and irregularly scalloped. Apical ocellus is entirely contained within M1-M2, round, black, ringed in rust orange, and with a single small white pupil at its center.

Hindwing ( Fig. 12 C, D): Wing rectangular, termen slightly convex and entire. Dorsal side similar in color to the forewing with the postmedian band visible as an obscure rust orange patch between M3-CuA2. Fringe scales are as in the forewing and long piliform scales appear at the base and over the discal cell, extending to the median and toward the inner margin. Ventral side similar in color to the dorsal side. Postmedian band lighter brown than the rest of the wing with both proximal and distal edges dark chocolate to dark coffee brown and scalloped.

Male genitalia ( Fig. 21 D–F): Uncus widest at the base, narrowing to an acute terminus and a little longer than the tegumen. Gnathos acute and almost three-fifths the length of the uncus. Pedunculus deltoid and wide. Saccus U-shaped and about equal in length to the gnathos. Valvae widest at the median, narrowing proximally to an acute triangle and narrowing slightly toward the distal end and a deltoid terminus. Aedeagus widest and truncate at the proximal end, narrowing gradual toward the distal end.

Specimens examined. Bolivia, Potosí Department, (CUIC) 1 male, 1 female

Figures

FIGURE 12. Tetraphlebia species. For composite habitus images, the left side is the dorsal surface and the right side is the ventral surgace. A, B: Tetraphlebia germainii C. Felder & R. Felder, 1867 male (BMNH # 809472) (A), female (MZUJ) (B); C, D: Tetraphlebia eleates (Weymer, 1890) n. comb., n. stat. male (C) and female (D) (CUIC); E: Tetraphlebia leucoglene (C. Felder & R. Felder, 1867) n. comb. male holotype (BMNH # 809770); F: Tetraphlebia patagonica n. comb., Miradores de Darwin, Ría Deseado, Santa Cruz, Argentina, 16 Nov. 2012 (photo by Carlos Agulian http: // www. ecoregistros. org / site _ en / imagen. php? id = 17587; used with permission).

FIGURE 21. Male genitalia of Tetraphlebia species. See Fig. 2 for labels of parts. All drawings to the same scale. A – C: Tetraphlebia germainii C. Felder & R. Felder, 1867 (CUIC), (A) showing uncus bent downwards; D – F: Tetraphlebia eleates (Weymer, 1890) n. comb. (CUIC); G – I: Tetraphlebia leucoglene (C. Felder & R. Felder, 1867) n. comb. (CH 30 - 1).