Neomaenas poliozona eustephanos Matz

Matz, Jess & Brower, Andrew V. Z., 2016, The South Temperate Pronophilina (Lepidoptera: Nymphalidae: Satyrinae): a phylogenetic hypothesis, redescriptions and revisionary notes, Zootaxa 4125 (1), pp. 1-108 : 42-43

Neomaenas poliozona eustephanos Matz stat. nov., nom. nov.

( Figs. 7 F; 26)

(objective replacement name for Neomaenas poliozona reedii ( Butler, 1881) (Neosatyrus) under ICZN Art. 60.3 (1999)). Holotype: (male) BMNH Type No. Rh3872

Type Location: Chile

Distribution. Found in Chile from the southwestern part of Bío-Bío province to Araucanía province at 50–200m from January to early March ( Fig. 26). Though the range of N. poliozona eustephanos is similar to that of N. poliozona poliozona , populations are allopatric. Ecological factors that separate these two subspecies are unknown and in need of further study.

Diagnosis. Identical to N. poliozona poliozona , but lacking the black spots on the ventral side of the hindwing, darker in color, and having a ripple pattern in dark chocolate striations over the hindwing.

Redescription. Head: Palps with a longitudinal black stripe bordered dorsally with a cream to white stripe. Dorsal side piliform scales on the palps are black and scales on the ventral side are black and tan.

Forewing ( Fig. 7 F): Dorsal side chocolate to dark chocolate brown, females with a patch of rust orange that extends from the discal cell to subterminal band. Postmedian band appears on the dorsal side of some males as a pair of rectangular patches between M3 and CuA2. Apical ocellus between M1-M3 appearing in some specimens as a round black spot. Ventral side similar in color to the dorsal side with a rust orange patch extending from the discal cell to the subterminal band, the postmedian band slightly lighter and bordered in dark chocolate. A ripple pattern with dark chocolate to dark coffee striations that extend along the costa and subterminal band.

Hindwing ( Fig. 7 F): Dorsal side similar in color to the forewing, postmedian band appearing in rust orange from M1 to the inner margin in females and rust red to rust orange in patches from M2 to CuA1 and from CuA2 to the inner margin. Postmedian band may not appear in darker specimens. Ventral side dark taupe to warm medium brown with a ripple pattern appearing in dark chocolate to dark coffee over the entire wing. Postmedian band is a saturated lavender, edged on the median side in dark chocolate. Hindwing ocelli rarely appear as small, round, black spots between Rs-M1, M1-M2, or CuA1-CuA2.

Remarks. The taxon bearing the name Neosatyrus reedii Butler, 1881 has been regarded previously as a separate species in a separate genus from N. poliozona poliozona by its original author and by Weymer (1911), and synonymized with N. poliozona by Ureta (1956) and Lamas & Viloria (2004). Herrera (1966) placed reedii in Neomaenas and noted its similarity to poliozona in genitalic features, also but differences in distribution and wing pattern, remarking that it may be a legitimate subspecies of N. poliozona .

Our decision to synonymize Spinantenna Hayward, 1953 , with Neomaenas Wallengren, 1858 , means that Neomaenas reedii ( Butler, 1881) (now viewed as a subspecies of N. poliozona ) becomes a secondary junior homonym of Neomaenas reedi ( Reed, 1877) , under Arts. 57.3.1 and 58.14 of the ICZN Code (ICZN 1999), because the latter is has priority as an available name, even though it is considered a subjective junior synonym of N. tristis ( Guerin-Ménéville, [1830]).

Etymology. the name refers to the purple submarginal band on the HWV.

Specimens examined. Chile, Araucanía Province, (MTSU) CL0518-CL0521, CL0601-0627, CL0823-0829; Chile, unknown province (BMNH) Holotype male

Figures

FIGURE 7. Composite habitus images of Neomaenas species. The left side of each specimen is the dorsal surface, the right side is the ventral surface. A, B: Neomaenas inornata Elwes 1903. Lithographs of male holotype (BMNH # Rh 3864) (A), and female allotype (BMNH) (B), (reproduced from Plate XIV of Elwes, 1903, Figure 19); C, D: Neomaenas monachus (Blanchard, 1852) — male N. monachus monachus (C), male N. monachus limonias (MZUJ) (D); E, F: Neomaenas poliozona (C. Felder & R. Felder, 1867) male holotype N. poliozona (BMNH # 809789) (E), male holotype N. poliozona eustephanos Matz nom. nov. (BMNH # 809619) (F); G: Neomaenas simplex (Butler, 1881) male holotype (BMNH # Rh 3878); H: Neomaenas tristis (Guérin-Méneville, [1830]) male (MZUJ); I, J: Neomaenas wallengrenii Butler, 1881, male lectotype (BMNH # 809361) (I), female paralectotype (BMNH # 809362) (J).