Huia, YANG, 1991

Arifin, Umilaela, Chan, Kin Onn, Smart, Utpal, Hertwig, Stefan T., Smith, Eric N., Iskandar, Djoko T. & Haas, Alexander, 2021, Revisiting the phylogenetic predicament of the genus Huia (Amphibia: Ranidae) using molecular data and tadpole morphology, Zoological Journal of the Linnean Society 193, pp. 673-699 : 687

GENUS HUIA YANG, 1991

( FIG. 6A)

Type species: Rana cavitympanum Boulenger, 1893 , Proceedings of the Zoological Society of London: 525. Holotype: BMNH 1893.5 .30.26 (BMNH 1947.2.4.16); adult male).

Type locality: Kina Baloo , North Borneo ( Kinabalu , Sabah, Malaysia) . Collector: A. Everett.

Common name: Sabah Huia frog (English) following Frank & Ramus (1995) and Frost (2019); Kongkang Jeram Sabah (Bahasa Indonesia).

Phylogenetic definition and content: Huia is a distinct and independently evolving genus, which shares a most recent common ancestor with the genus Meristogenys ( Fig. 2). It is currently understood to be monotypic, containing H. cavitympanum , which is endemic to the island of Borneo.

Diagnosis: (Tadpole characters presented here are based on observation by U.A., whereas morphological characters of adults are taken from published studies (e.g. Inger, 1966; Yang, 1991; Manthey & Denzer, 2014). Larvae gastromyzophorous; upper beak Λ- shaped, lower beak V-shaped, both undivided; LTRF 10–12((3– 4)–(10–12))/5–6(0–2); postorbital glands present; an oblique cluster of glands immediately below the spiracular tube, running upward and backward present; prespiracular glands present in some individuals ( Inger, 1985); glands on the tail fin absent. Adults medium in size, body stocky; snout rounded; tympanum clear, deep in cavity; supra- and posttympanic folds present, dorsolateral folds present; ratio of tibia length to snout–vent length> 70%; outer metatarsal tubercle present or absent (absent according to Yang, 1991), inner metatarsal tubercle present, small; length of Finger I = Finger II; length of metacarpal of Finger I 54% of Finger II; width of crossbar terminal phalanx of Finger III 50% length; male with nuptial pad and paired lateral vocal sacs; humeral gland absent. In life, dorsum generally dark brown; flanks light brown with markings; head light brown with scattered dark brown markings; lips with dark markings.