Clubiona, Wagner, 1887

Bosmans, Robert, Henrard, Arnaud, Benhalima, Souâd & Kherbouche-Abrous, Ourida, 2017, The genus Clubiona Latreille, 1904 (Araneae: Clubionidae) in the Maghreb, with notes on the genevensis group and new records from the Mediterranean Region, Zootaxa 4353 (1), pp. 1-28: 3-4

publication ID

https://doi.org/10.11646/zootaxa.4353.1.1

publication LSID

lsid:zoobank.org:pub:7D342E35-8D42-4C3D-9903-49E5F08D7D34

persistent identifier

http://treatment.plazi.org/id/03EF9555-0904-FF99-1DD1-FA781790FB83

treatment provided by

Plazi

scientific name

Clubiona
status

 

Key to the Clubiona   species of the genevensis   group

Clubiona minor   , C. pseudominor   and C. wunderlichi   were not examined and, for these species, the key is based on characters found in literature.

1 Males (those of C. pseudominor   and C. wunderlichi   unknown).................................................. 2

- Females............................................................................................. 7

2(1) Embolus originating in distal or median part of bulbus and directed anteriorly ( Figs 4 View FIGURES 1–8 , 12 View FIGURES 9–15 , 17 View FIGURES 16–19 ; Wunderlich 1987: fig. 640), basolateral extension of bulbus and cymbium short, as long as wide ( Figs 54, 55 View FIGURES 52–67 ); modified cymbial setae long: as long as half the length of cymbium or more ( Figs 3, 4 View FIGURES 1–8 , 11 View FIGURES 9–15 , 16 View FIGURES 16–19 , 52 View FIGURES 52–67 )..........................................3 ( decora   subgroup)

- Embolus originating more basally and directed laterally ( Figs 25 View FIGURES 20–25 , 38 View FIGURES34–41 , 46 View FIGURES 45–51 ); basolateral extension of bulbus and cymbium much longer than wide ( Figs 59, 63, 67 View FIGURES 52–67 ); modified cymbial setae shorter, length less than half length of cymbium ( Figs 24 View FIGURES 20–25 , 37 View FIGURES34–41 , 45 View FIGURES 45–51 , 56, 60, 64 View FIGURES 52–67 )5........................................................................ ( genevensis   subgroup)

3(2) Palpal tibia nearly twice as long as wide ( Fig. 11 View FIGURES 9–15 ); RTA tip more rounded ( Fig. 11 View FIGURES 9–15 ); origin of embolus in distal part of bulbus ( Figs 12 View FIGURES 9–15 , 17 View FIGURES 16–19 )................................................................................ C. diniensis  

- Palpal tibia slightly longer than wide ( Fig. 3 View FIGURES 1–8 ; Wunderlich 1987: fig. 641); RTA with sharp tip ( Fig. 3 View FIGURES 1–8 ; Wunderlich 1987: fig. 641); origin of embolus in median part of bulbus ( Fig. 4 View FIGURES 1–8 ; Wunderlich 1987: fig. 640)................................ 4

4(3) RTA thin, triangular, not broadened basally ( Fig. 3 View FIGURES 1–8 ).................................................... C. decora  

- RTA with broad base extending dorsally ( Wunderlich 1987: figs 641, 642).................................. C. minor  

5(2) Retrolateral tibial apophysis (RTA) rounded, slightly longer than wide at its base ( Figs 37 View FIGURES34–41 , 62 View FIGURES 52–67 ); ventral profile of anterior part of bulbus concave ( Figs 37 View FIGURES34–41 , 60 View FIGURES 52–67 ); modified cymbial setae with spherical sub-basal swelling ( Figs 74–76 View FIGURES68–79 ); abdomen dorsally with pale spot on posterior half ( Fig. 34 View FIGURES34–41 ).......................................................... C. leucaspis  

- RTA roughly triangular or bluntly pointed ( Figs 24 View FIGURES 20–25 , 45 View FIGURES 45–51 , 57, 58, 65, 66 View FIGURES 52–67 ); ventral profile of bulbus convex ( Figs 24 View FIGURES 20–25 , 45 View FIGURES 45–51 , 56, 64 View FIGURES 52–67 ); modified cymbial setae with flattened sub-basal swelling ( Figs 73, 79 View FIGURES68–79 ); abdomen without postero-dorsal white spot ( Figs 20– 23 View FIGURES 20–25 , 42, 43 View FIGURES 42–44 )........................................................................................... 6

6(5) Chelicerae not enlarged ( Figs 20, 22, 23 View FIGURES 20–25 ); RTA roughly triangular, with blunt, rounded tip ( Figs 24 View FIGURES 20–25 , 57, 58 View FIGURES 52–67 ); modified cymbial setae with well developed, plate shaped sub-basal swelling ( Figs 72, 73 View FIGURES68–79 )............................... C. genevensis  

- Chelicerae dark brown, strongly protruding ( Figs 43, 44 View FIGURES 42–44 ); RTA triangular with sharper tip ( Figs 45 View FIGURES 45–51 , 65, 66 View FIGURES 52–67 ); sub-basal swelling of modified cymbial setae reduced, crescent shaped ( Fig. 79 View FIGURES68–79 )............................................ C. vegeta  

7(1) Copulatory ducts loose, slightly coiled or not, connected laterally to the atrio-spermathecal part ( Figs 6–8 View FIGURES 1–8 , 13–15 View FIGURES 9–15 ; Wunderlich 1987: figs 643–645; Mikhailov 1992: figs 2A, B)............................................ 8 ( decora   subgroup)

- Copulatory ducts tightened, strongly convoluted, connected inferiorly to the atrio-spermathecal part ( Figs 29–32 View FIGURES 26–33 , 40, 41 View FIGURES34–41 , 48– 51 View FIGURES 45–51 )............................................................................. 12 ( genevensis   subgroup)

8(7) Copulatory ducts straight forward, not coiled ( Wunderlich 1987: figs 643–645, 645a)..................................................................................................................................... 9

- Copulatory ducts coiled ( Figs 6–8 View FIGURES 1–8 , 13–15 View FIGURES 9–15 ; Mikhailov 1992: figs 2A, B).......................................... 10

9(8) Fertilization duct well developed, conical ( Wunderlich 1987: fig. 644)...................................... C. minor  

- Fertilization duct thinner, tube-shaped ( Wunderlich 1987: fig. 645a)................................. C. pseudominor  

10(8) Epigyne with two distinct oval atria ( Figs 13, 14 View FIGURES 9–15 , 18 View FIGURES 16–19 )................................................ C. diniensis  

- Epigyne with only one, roughly inverted heart-shaped atrium ( Figs 6–8 View FIGURES 1–8 ; Mikhailov 1992: figs 2A, B).................. 11

11(10) Spermathecae far away from the anterior part of longitudinal copulatory ducts ( Figs 6– 8 View FIGURES 1–8 )..................... C. decora  

- Anterior part of longitudinal copulatory ducts almost reaching the inferior part of spermathecae ( Mikhailov 1992: figs 2A, B)......................................................................................... C. wunderlichi  

12(7) Abdomen dorsally with short anterior median stripe and with contrasting pale oval area on posterior half ( Figs 35, 36 View FIGURES34–41 ); spermathecae small, separated by their diameter or more ( Figs 39–41 View FIGURES34–41 )...................................... C. leucaspis  

- Abdomen usually with longer, sometimes interrupted median stripe, posteriorly without dorsal pale spot ( Figs 21 View FIGURES 20–25 , 42 View FIGURES 42–44 ); spermathecae larger, separated by less than their diameter ( Figs 26–33 View FIGURES 26–33 , 47–51 View FIGURES 45–51 )....................................... 13

13(12) Dorsal median stripe most often absent in anterior third of abdomen, fading into V-shaped stripes in posterior third ( Fig. 21 View FIGURES 20–25 ); copulatory opening often narrower, longer than wide ( Figs 26, 27 View FIGURES 26–33 ); vulva with narrower atrium ( Figs 30, 32 View FIGURES 26–33 )... C. genevensis  

- Dorsal median stripe often reaching anterior margin of abdomen, tapered posteriorly and fading in posterior third ( Fig. 42 View FIGURES 42–44 ; better preserved in the male Fig. 43 View FIGURES 42–44 ); copulatory opening wider, at least as wide as long ( Figs 47, 48 View FIGURES 45–51 ); vulva with wider atrium ( Figs 49, 51 View FIGURES 45–51 ).................................................................................. C. vegeta