Grapsus fourmanoiri Crosnier, 1965

Ng, Ngan Kee, Rodríguez Moreno, Paula A., Naruse, Tohru, Guinot, Danièle & Mollaret, Noémy, 2019, Annotated type-catalogue of Brachyura (Crustacea, Decapoda) of the Muséum national d’Histoire naturelle, Paris. Part II. Gecarcinidae and Grapsidae (Thoracotremata, Grapsoidea), with an Appendix of pre- 1900 collectors, Zoosystema 41 (7), pp. 91-130 : 104

Grapsus fourmanoiri Crosnier, 1965

( Fig. 3E )

Grapsus fourmanoiri Crosnier, 1965: 12-17 , figs 4-6, pl. 3, fig. 1, table p. 16.

CURRENT TAXONOMIC STATUS. — Grapsus fourmanoiri Crosnier, 1965 .

LECTOTYPE (by present designation). — MNHN-IU-2008-10635 (= MNHN-B11803 ), ♂ 28.0 × 31.7 mm, Nosy Bé, Madagascar, coll. and det. A. Crosnier.

PRESERVATION OF THE LECTOTYPE. — In alcohol. RP1 missing.

PARALECTOTYPES. — MNHN-IU-2014-11212 (= MNHN-B11531), 3♂ 22.6 × 25.2 – 24.2 × 27.8 mm, 1 ♀ 21.2 × 24.4 mm, 2 ovig. ♀ 24.1 × 28.6 – 29.5 × 33.4 mm, same data as lectotype .

PRESERVATION OF THE PARALECTOTYPES. — In alcohol.

REMARKS

Crosnier (1965: 14, table p. 16) has based the species on several specimens from Nosy Bé, Madagascar, but he did not expressly mention any type specimens in his publication. He mentions in the text a large specimen (length: 34.1; width: 37.7 mm) and provides measurements of five males and two females in a table ( Crosnier 1965: 16). Two lots of specimens deposited at the Muséum, MNHN-IU-2008-10635 (= MNHN-B11803) and MNHN-IU-2014-11212 (= MNHN-B11531) are labelled “ Syntypes ”. Our measurements of most of the specimens do not correspond to the dimensions provided by Crosnier (1965: 16), except for one male MNHN-IU-2008-10635 (= MNHN-B11803), 28.0 × 31.7 mm. This discrepancy can be easily accounted for by how the specimens were measured. The largest specimen (length: 34.1; width: 37.7 mm) could not be located. The next largest male specimen MNHN-IU-2008-10635 (= MNHN-B11803) is here designated as lectotype of G. fourmanoiri . The other specimens from the type series become paralectotypes. It is very likely that Crosnier (1965) has collected and examined more than eight specimens belonging to his type series. Actually, one specimen was donated to Leiden Museum by Crosnier in 1966 (RMNH D 22683, male 23.7 × 27.0 mm, Nosy Bé, Madagascar, coll. and det. A. Crosnier): it is indicated as syntype in the Catalogue of the Types of the Nationaal Natuurhistorisch Museum, Leiden, by Fransen et al. (1997: 123) and shows the same ‘type and form’ of label as the labels of the MNHN jars. This RMNH sample also becomes paralectotype.

Fransen et al. (1997: 123, footnote) regarded two additional samples as syntypes of G. fourmanoiri , but this mention corresponds to an erroneous statement. Crosnier (1965: 17, footnote) wrote: “At Leiden, we have reexamined specimens from Banerjee (1960: 149) from Durban and Reunion Rocks, initially identified as Grapsus albolineatus ”. According to Fransen (pers. comm., December 2012), two samples (RMNH D3210, one female 32.5 × 35.0 mm, South Africa, Reunion Rocks, 28.X.1938, leg. L. D. Brongersma; RMNH D3250, 2 males 10.8-16.9 × 13.0- 19.6 mm, South Africa, Durban, 10-11.XI.1938, leg. H. Engel) have been deemed by Crosnier as suitable syntypes of his G. fourmanoiri . However, despite the Code ’s Recommendation (Art. 74E) about the verification of locality (“ When selecting a lectotype, the author should, if possible, verify the accuracy of the locality ascribed to it”), it is obvious that these two samples, RMNH D3250 from Durban and RMNH D3210 from Reunion Rocks, have not been collected from Nosy Bé, Madagascar by A. Crosnier. It is likely that they were only subsequently identified by him as G. fourmanoiri and, therefore, they cannot be considered syntypes, even if they correspond to G. fourmanoiri .

Figures

FIG. 3. — A, Pelocarcinus marchei A. Milne-Edwards, 1890, MNHN-IU-2000-3752 (= MNHN-B3752), lectotype, ♂; B, Thelphusa rotunda Quoy & Gaimard, 1824, MNHN-IU-2000-3745 (= MNHN-B3745), lectotype, ♂; C, Grapsus brevipes H. Milne Edwards, 1853, MNHN-IU-2000-10978 (= MNHN-B10978), lectotype, ♂; D, Grapsus cruentatus Latreille, 1803, MNHN-IU-2000-3407 (= MNHN-B3407), neotype, ♂; E, Grapsus fourmanoiri Crosnier, 1965, MNHN-IU-2008-10635 (= MNHN-B11803), lectotype, ♂; F, Grapsus gracilipes H. Milne Edwards, 1853, MNHN-IU-2000-3411 (= MNHN-B3411), lectotype, ♂; G, Grapsus granulosus H. Milne Edwards,1853,MNHN-IU-2000-3417 (= MNHN-B3417),lectotype,♂; H, Grapsus grayi H. Milne Edwards,1853,MNHN-IU-2000-3421 (= MNHN-B3421), lectotype, ♀. Scale bars: A-D, F, H, 2 cm; E, 10 mm; G, 5 cm.