Cephennodes (Cephennodes) mangmayanus, Jałoszyński, 2017

Jałoszyński, Paweł, 2017, Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 4349 (1), pp. 1-120 : 74-75

Cephennodes (Cephennodes) mangmayanus sp. n.

( Figs 161 , 215–218 , 229 )

Type material. Holotype: NEPAL ( Sankhuwasabha District ): ♂, one label: "E. NEPAL: KOSI / Crête N-E Mangmaya / 2800 m, 7.IV.84 / Löbl - Smetana" [white, printed] ( MHNG) . Paratypes (2 exx): 1 ♀, same data as for holotype ; 1 ♂, "E. NEPAL: KOSI / Forêt S. Mangsingma / 2300 m, 13.IV.84 / Löbl - Smetana" [white, printed] ( MHNG, cPJ).

Diagnosis. BL 1.33–1.43 mm; abdominal sternite II with posterior microserrate marginal carina which is shallowly but distinctly emarginate at middle and indistinctly at each side, sternite III with much shorter microserrate lamina slightly expanded posteriorly and broadly rounded, lamina strongly projected ventrally, posterior margin of sternite III slightly expanded posteriorly, broadly rounded; apex of median lobe of aedeagus subtrapezoidal, blunt and distinctly asymmetrical; apical projection of aedeagus distant from apex of median lobe, with distal margin convex proximally and shallowly concave in median region, strongly oblique in relation to the long axis of aedeagus, apical portion of apical projection subtriangular, nearly pointed.

Description. Body of male ( Fig. 161 ) dark brown, moderately stout, with shallow constriction between pronotum and elytra, strongly convex, covered with light brown setae; BL 1.33–1.43 mm.

Head subtrapezoidal, HL 0.18 mm, HW 0.40 mm; vertex and frons confluent and weakly, evenly convex, vertex with a pair of tiny but distinct tubercles; supraantennal tubercles small, weakly elevated; eyes moderately large, strongly convex, coarsely faceted. Punctures on vertex and frons small and shallow but distinct, moderately sharply marked, nearly evenly and densely distributed, separated by spaces slightly shorter than diameters of punctures; setae sparse, short, suberect. Antennae moderately long and moderately thickened, AnL 0.73–0.75 mm, pentamerous club indistinctly delimited; antennomere I about 1.3 × as long as broad; II narrower but slightly longer than I, 1.5 × as long as broad; III–VI each slightly narrower and much shorter than II, about as long as broad to 1.1 × as long as broad; VII indistinctly broader and indistinctly longer than VI, about 1.2 × as long as broad; VIII distinctly broader and slightly shorter than VII, about as long as broad; IX and X each distinctly larger than preceding ones, each slightly transverse; XI distinctly broader than X, as long as IX–X together, about 1.6 × as long as broad.

Pronotum in dorsal view subtrapezoidal, broadest shortly in front of base but very indistinctly narrowing both posteriorly and anteriorly up to anterior third, PL 0.40–0.43 mm, PW 0.68–0.70 mm; anterior margin in strictly dorsal view straight; lateral margins indistinctly microserrate, strongly rounded in anterior third, nearly straight in posterior half; hind corners nearly right-angled; posterior margin shallowly bisinuate; lateral carinae distinct but narrow, fused with lateral margins; antebasal pits small and shallow, each much closer to posterior than to lateral pronotal margin. Punctures on disc similarly distinct and dense as those on frons and vertex; setae moderately long, dense, suberect.

Elytra elongate, oval, broadest slightly in front of middle, EL 0.75–0.78 mm, EW 0.70 mm, EI 1.07–1.11. Humeral calli weakly elevated; subhumeral lines distinct, divergent posteriorly, equal to about 0.3 EL, each developed as moderately sharp stepwise border between more convex humeral region and less convex adsutural area; basal pit on each elytron connected to shallow and diffuse arcuate impression extending posteromesally; apices of elytra separately rounded. Punctures slightly larger but more diffuse than those on pronotum, similarly dense; setae similar to those on pronotum.

Hind wings not studied.

Metaventrite with moderately large, sharply delimited postmesocoxal impressions, with median area covered with fine, dense punctures.

Abdomen ( Figs 215–216 ) strongly modified; sternite II with posterior microserrate marginal carina which is shallowly but distinctly emarginate at middle and indistinctly at each side, sternite III with much shorter microserrate lamina slightly expanded posteriorly and broadly rounded, lamina strongly projecting ventrally, in lateral view ( Fig. 215 ) visible as a narrow and long projection; posterior margin of sternite III slightly expanded posteriorly, broadly rounded.

Legs unmodified.

Aedeagus ( Figs 217–218 ) moderately slender, AeL 0.20 mm, apex of median lobe subtrapezoidal and distinctly asymmetrical; apical projection distant from apex of median lobe, its distal margin convex proximally and weakly concave in median region, so strongly oblique in relation to the long axis of aedeagus that it becomes lateral margin; apex of apical projection subtriangular and nearly pointed; parameres very slender, slightly unequal in length, one reaching and one not reaching apex of median lobe, each with one subapical seta.

Female. Similar to male but with unmodified abdomen; BL 1.43 mm; HL 0.18 mm, HW 0.40 mm, AnL 0.73 mm; PL 0.45 mm, PW 0.70 mm; EL 0.80 mm, EW 0.73 mm, EI 1.10.

Distribution ( Fig. 229 ). Eastern Nepal.

Etymology. Locotypical, after Mangmaya, a locality east of Makalu Barun National Park.

Remarks. This species can be distinguished from all remaining members of the C. simplicipes group on the basis of the abdominal modifications in lateral view: the lamina on sternite III is very long, narrow and strongly projected ventrally.

Figures

FIGURES 156 – 162. Dorsal habitus of Cephennodes (s. str.) paralalitpuranus sp. n. (156), C. (s. str.) bahrabisensis sp. n. (157), C. (s. str.) bilaminatus sp. n. (158), C. (s. str.) ghorepanianus sp. n. (159), C. (s. str.) cordilaminatus sp. n. (160), C. (s. str.) mangmayanus sp. n. (161), and C. (s. str.) bilobatus sp. n. (162).

FIGURES 211 – 222. Abdomen of male in right lateral (211, 215, 219) and ventral (212, 216, 220) views and aedeagus in ventral (213, 217, 221) and lateral (214, 218, 222) views. Cephennodes (s. str.) cordilaminatus sp. n. (211 – 214), C. (s. str.) mangmayanus sp. n. (215 – 218), and C. (s. str.) bilobatus sp. n. (219 – 222).

FIGURES 229 – 230. Distribution of Cephennodes (s. str.) cordilaminatus sp. n. (229, circle), C. (s. str.) mangmayanus sp. n. (229, triangle), C. (s. str.) bilobatus sp. n. (230, circle).