Cephennodes (Cephennodes) pampinosus, Jałoszyński, 2017

Jałoszyński, Paweł, 2017, Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 4349 (1), pp. 1-120 : 16-17

Cephennodes (Cephennodes) pampinosus sp. n.

( Figs 20 , 24–26, 29–30 , 31 )

Type material. Holotype: NEPAL (Lalitpur District): ♂, one label: " NEPAL (Prov. Bagmati) / Phulchauki nr. Kath- / mandu, 1700 m, 10.V.81 / I. Löbl" [white, printed] ( MHNG). Paratypes (2 exx): 2 ♀♀, same data as for holotype ( MHNG and cPJ).

Diagnosis. Head lacking impression; frons between eyes with median longitudinal carina; setae on frons in front of eyes modified, hyaline and leaf-shaped.

Description. Body of male ( Fig. 20 ) dark brown, very stout, with shallow constriction between pronotum and elytra, moderately convex, covered with light brown setae; BL 1.38–1.40 mm.

Head ( Figs 24–26 ) subtrapezoidal, HL 0.23 mm, HW 0.34–0.35 mm; posterior portion of vertex convex, anterior portion of vertex and most of frons flattened, with short median longitudinal carina, which is angulate in lateral view ( Fig. 25 ); supraantennal tubercles distinct but weakly elevated; eyes moderately large, strongly convex, coarsely faceted. Punctures on posterior portion of vertex inconspicuous, those on its flattened anterior area and on median region of frons small but sharply marked and very dense, separated by spaces distinctly shorter than diameters of punctures; setae on vertex and anterior margin of clypeus thin, long and suberect, those on flattened median area of frons strongly modified, leaf-like and hyaline, additionally median carina flanked by two median brushes of thin setae directed laterally and two posterior brushes directed anteriorly. Antennae long and slender, moderately strongly thickened, AnL 0.75 mm; antennomeres relatively loosely assembled, pentamerous club indistinctly delimited and about as long as half AnL; antennomere I about 1.7 × as long as broad; II slightly narrower and shorter than I, 1.5 × as long as broad; III and IV subequal in shape, each narrower than II and subquadrate; V distinctly longer than IV but indistinctly broader, 1.5 × as long as broad, VI slightly shorter and narrower than V, 1.5 × as long as broad; VII distinctly broader than VI but indistinctly longer, slightly elongate; VIII about as broad as VII but indistinctly shorter, slightly elongate, IX distinctly broader than VIII but indistinctly longer, about as broad as long, X distinctly broader and longer than IX, about as long as broad, XI only slightly broader than X, about as long as IX–X together, nearly twice as long as broad.

Pronotum in dorsal view subtrapezoidal, broadest at base, PL 0.40–0.43 mm, PW 0.65 mm; anterior margin in strictly dorsal view slightly concave; lateral margins indistinctly microserrate, strongly rounded in anterior half and weakly so in posterior third; hind pronotal corners nearly right-angled; posterior margin shallowly bisinuate; lateral carinae narrow, fused with lateral margins; antebasal pits small and shallow, diffuse, each located much closer to posterior than to lateral pronotal margin. Punctures on disc very small but deep and sharply marked, unevenly distributed, those on median area separated by spaces 0.5–1 × as wide as diameters of punctures; setae long, moderately dense, suberect.

Elytra very stout, oval, broadest in front of middle, EL 0.75 mm, EW 0.75 mm, EI 1.00. Humeral calli weakly elevated; subhumeral lines nearly as long as half EL, narrow and sharply marked, developed as sharp stepwise border between more convex humeral region and less convex adsutural area; basal pit on each elytron large and located in short, deep, nearly round basal impression; apices of elytra separately rounded. Punctures slightly larger and sparser than those on pronotum; setae similar to those on pronotum.

Hind wings not studied.

Metaventrite with very large, distinctly delimited postmesocoxal impressions exceeding half length of ventrite, median area covered with fine punctures.

Abdomen and legs unmodified.

Aedeagus ( Figs 28–29 ) in ventral view nearly symmetrical, AeL 0.21 mm, median lobe oval with subtriangular narrow apex; apical projection in lateral view hook-like, strongly curved dorsobasally; parameres slender, slightly unequal in length, each with three long thin setae in apical and subapical region.

Female. Very similar to male but with unmodified frons covered with thin setae; BL 1.40 mm; HL 0.23 mm, HW 0.35 mm, AnL 0.75 mm; PL 0.43 mm, PW 0.65 mm; EL 0.75 mm, EW 0.75 mm, EI 1.00.

Distribution ( Fig. 31 ). Eastern Nepal.

Etymology. The name pampinosus (Latin leafy, covered with leaves) refers to the modified setae on the vertex.

Remarks. Strongly modified, hyaline leaf-like setae covering the frons in C. pampinosus are unique, no other species of Cephennodes has such setae, only in some Leptomastacini setae are broad and flattened, but not hyaline.

Figures

FIGURE 31. Distribution of Cephennodes (s. str.) cavifrons sp. n. (circle) and C. (s. str.) pampinosus sp. n. (triangle).

FIGURES 19 – 20. Dorsal habitus of Cephennodes (s. str.) cavifrons sp. n. (19) and C. (s. str.) pampinosus sp. n. (20).

FIGURES 21 – 30. Head in anterior (21, 24), dorsolateral (22, 25) and dorsal (23, 26) views, aedeagus on ventral (27, 29) and lateral (28, 30) views. Cephennodes (s. str.) cavifrons sp. n. (21 – 23, 27 – 28) and C. (s. str.) pampinosus sp. n. (24 – 26, 29 – 30).