Cephennodes (Cephennodes) manangensis, Jałoszyński, 2017

Jałoszyński, Paweł, 2017, Ant-like stone beetles on the roof of the world. Cephenniini of Nepal and Bhutan (Coleoptera, Staphylinidae, Scydmaeninae), Zootaxa 4349 (1), pp. 1-120 : 41-43

Cephennodes (Cephennodes) manangensis sp. n.

( Figs 95 , 103, 105 , 116–119 , 144 )

Type material. Holotype: NEPAL (Manang District): ♂, one label: " NEPAL, Manang Distr. / above Bagarchap, 2200 m / 12.–13.IV.1980 / leg. J. Martens & A. Ausobsky" [white, printed] ( SMNS). Paratype: 1 ♂, same data as for holotype ( SMNS).

Diagnosis. BL 1.25–1.30 mm; body relatively slender; distal portion of metatibia strongly thickened, with oval subapical porous field much longer than broad and lacking one long bristle within short setal fringe of apical margin; abdomen in ventral view with sternite III bearing broadly rounded median lobe; apical portion of median lobe of aedeagus evenly rounded, its longitudinal axis forming about 45°-angle with long axis of aedeagus; apical projection with sinuate distal margin.

Description. Body of male ( Fig. 95 ) moderately dark brown, with shallow constriction between pronotum and elytra, moderately convex, covered with light brown setae; BL 1.25–1.30 mm (mean 1.28 mm).

Head subtrapezoidal, HL 0.18 mm, HW 0.18–0.20 mm (mean 0.19 mm); vertex broadly rounded, with a pair of tiny tubercles; frons flattened; supraantennal tubercles distinct but weakly elevated; eyes large and strongly convex, coarsely faceted. Punctures on vertex and frons distinct, small but sharply marked and densely, evenly distributed, separated by spaces as wide as 1–2 × diameters of puncture; setae short, moderately dense, suberect. Antennae short, AnL 0.63 mm, compact, pentamerous club indistinctly delimited and moderately strongly thickened; antennomere I twice as long as broad; II slightly shorter and narrower than I, 1.8 × as long as broad; III– IV each slightly narrower than II, subquadrate; V as broad as IV but slightly longer, 1.2 × as long as broad; VI as broad as V but slightly shorter, subquadrate; VII slightly broader and longer than VI, as long as broad; VIII slightly broader but minimally shorter than VII, slightly transverse; IX much larger than VIII, as long as broad; X distinctly larger than IX, slightly broader than long; XI distinctly broader than X, slightly longer than IX–X together, about 2.2 × as long as broad.

Pronotum in dorsal view nearly semicircular, broadest slightly in front of middle, PL 0.38–0.40 mm (mean 0.39 mm), PW 0.58–0.60 mm (mean 0.59 mm); anterior margin broadly rounded; lateral margins indistinctly microserrate, rounded in anterior half, then nearly straight and slightly convergent toward obtuse-angled hind corners; posterior margin distinctly bisinuate; lateral carinae narrow, fused with lateral margins; antebasal pits small and shallow, each located much closer to posterior than to lateral pronotal margin. Punctures on disc much smaller and shallower than those on head, but very dense; setae dense, short, suberect.

Elytra short, oval, broadest distinctly anterior to middle, EL 0.70–0.73 mm (mean 0.71 mm), EW 0.63 mm, EI 1.12–1.16. Humeral calli weakly elevated but distinct; subhumeral lines moderately distinct, equal to about 0.3 EL, each developed as moderately sharply marked stepwise border between more convex humeral region and less convex adsutural area; basal pit on each elytron large and connected to short, elongate impression extending posteromesally; apices of elytra separately rounded. Punctures very small, sharply marked, dense, slightly more distinct than those on pronotum; setae short, dense, suberect.

Hind wings not studied.

Metaventrite with indistinctly delimited postmesocoxal impressions, remaining surface covered with very fine punctures, glossy.

Hind legs with strongly modified tibiae ( Figs 103, 105 ); in dorsal view lateral (external) tibial margin nearly straight in proximal half and evenly convex in distal half, spatulate portion of tibia gradually narrowed distally; dorsodistal porous field much longer than broad; distal fringe of setae lacking one long bristle.

Abdomen ( Figs 116–117 ) with sternites II and III distinctly projected in lateral view ( Fig. 116 ); in ventral view sternite III with broadly rounded median lobe ( Fig. 117 ).

Aedeagus ( Figs 118–119 ) moderately stout; AeL 0.18 mm, apical portion of median lobe evenly rounded, its longitudinal axis forming about 45°-angle with the long axis of aedeagus; apical projection with sinuate distal margin; parameres long and slender, not exceeding apex of median lobe, each bearing one apical and one subapical seta.

Female. Unknown.

Distribution ( Fig. 144 ). Central Nepal.

Etymology. Locotypical; after the Manang District in Nepal.

Remarks. This species can be identified on the basis of its small and slender body, the strongly elongate porous field on metatibia, the gradually narrowed spatulate portion of metatibia, and the moderately slender aedeagus with rounded apex of median lobe strongly curved outward from the long axis of aedeagus.

Figures

FIGURES 94 – 101. Dorsal habitus of Cephennodes (s. str.) dolakhanus s sp. n. (94), C. (s. str.) manangensis sp. n. (95), C. (s. str.) martensi sp. n. (96), C. (s. str.) paramartensi sp. n. (97), C. (s. str.) monolaminatus sp. n. (98), C. (s. str.) thakanus sp. n. (99), C. (s. str.) annapurnaensis sp. n. (100), and C. (s. str.) parbatensis sp. n. (101).

FIGURES 102 – 111. Right metatibia of male in dorsal view. Cephennodes (s. str.) dolakhanus s sp. n. (102, 104), C. (s. str.) manangensis sp. n. (103, 105), C. (s. str.) martensi sp. n. (106), C. (s. str.) paramartensi sp. n. (107), C. (s. str.) monolaminatus sp. n. (108), C. (s. str.) thakanus sp. n. (109), C. (s. str.) annapurnaensis sp. n. (110), and C. (s. str.) parbatensis sp. n. (111). 104 – 111 not to the same scale.

FIGURES 112 – 127. Abdomen of male in right lateral (112, 116, 120, 124) and ventral (113, 117, 121, 125) views and aedeagus in ventral (114, 118, 122, 126) and lateral (115, 119, 123, 127) views. Cephennodes (s. str.) dolakhanus sp. n. (112 – 115), C. (s. str.) manangensis sp. n. (116 – 119), C. (s. str.) martensi sp. n. (120 – 123), C. (s. str.) paramartensi sp. n. (124 – 127). Abbreviations: aml, apex of median lobe; ap, apical projection.

FIGURES 144 – 145. Distribution of Cephennodes (s. str.) dolakhanus sp. n. (144, circle), C. (s. str.) manangensis sp. n. (144, triangle), C. (s. str.) martensi sp. n. (145, circle), C. (s. str.) paramartensi sp. n. (145, triangle).