Calophya clavuligera, Burckhardt & Basset, 2000

Burckhardt, D. & Basset, Y., 2000, The jumping plant-lice (Hemiptera, Psylloidea) associated with Schinus (Anacardiaceae): systematics, biogeography and host plant relationships, Journal of Natural History 34 (1), pp. 57-155 : 84-85

Calophya clavuligera sp. n.

(®gure 13) Psyllid gall, RuÈ bsaamen, 1908: 77, ®gure 17. Brazil: (Parana Â?) Ipiranga, 21.iii.1905, galls on the leaves of Schinus sp. ([sic] probably L ithrea sp.) (Usteri) (depository?)

D escription . Adult unknown.

Fifth instar larva (®gure 13). Yellow, dorsum with brown median streak stretching from apex of head to abdominal tip. Eyes greyish, antennal apices dark brown. Dorsum ¯attened, disk-shaped, almost circular, sclerotized. Venter membraneous, strongly expanded to ®t into pit. Anterior margin of head almost straight, weakly, narrowly cleft in the middle. Antennae irregularly triangular, with small, slender sectasetae. Legs with very small claws. Anterior margin of humeral lobes ending distal to anterior eye margin. Abdominal dorsum without median row of horns. Caudal plate ratio 0.55. Circumanal ring oval. Head, wing bud and abdominal margins fringed with very slender sectasetae.

Host plants, gall and biology. L ithrea brasiliensis and molleoides . Inducing pit galls on the leaves; pit circular in outline with slightly swollen margins which are lighter than surrounding tissue. Galls and larvae have been described by RuÈ bsaamen (1908).

Distribution. Argentina (Catamarca, Cordoba, Corrientes, Misiones, Salta), Bolivia (Santa Cruz), Brazil (Rio Grande do Sul), Paraguay (Caaguazu Â, Canendiyu), Uruguay (Lavalleja, Rivera, Tacuarembo Â). The species was reported as unidenti®ed psyllid gall from Brazil (Parana Â?) (RuÈ bsaamen, 1908).

Material examined. HOLOTYPE ®fth instar larva, Argentina: Misiones Province , Rt. 14, 3 km E Centinela, 29.xi.1995, L ithrea molleoides (H. A. Cordo) , dry mounted ( NHMB).

PARATYPES. Argentina: 214 larvae, same data as holotype ( BMNH, NHMB, SABC, USNM) .

Material not included in type series. Argentina: pit galls on the leaves, Catamarca, Fuerte de Andalgala Â, x.1872, L ithrea molleoides (F. Schickendantz) ; deep pit galls on lower leaf surface, Cordoba, Diquede los Molinos, Sierra, 28.x.1974, L ithrea molleoides (Tirel) ; pit galls on the leaves, Cordoba, Ochoa, Punilla, 17.xi.1902 and 3.x.1904, L ithrea molleoides (T. Stuckert) ; larvae in pit galls on the leaves, Corrientes, Santo Tome Â, Ruta 40, Ayo. Chimiray, 12.xi.1974, L ithrea molleoides (A. Schinini and R. Carnevali) ; pit galls on the leaves, Corrientes, Ituzaingo Â, Isla Apipe  Grande, Puerta Mora, 11.xii.1973, L ithrea molleoides (A. Kapovickas et al.) ; leaves of L ithrea molleoides with pit galls and larvae, Misiones, same data as holotype; pit galls on the leaves, Misiones, Iguazu Â, Puerte Esperanza, 12 km, 16.x.1950, L ithrea molleoides (J. E. Montes) ; deep pit galls on the leaves, Salta, Guachipas, Cuesta del Lajar, along road, 1850 m, 27.x.1991, L ithrea molleoides (A. Charpin) . Bolivia: larvae in pit galls on the leaves, Santa Cruz, Samaipata, 19.x.1928, 1600 m, L ithrea molleoides , mountain forest (J. Steinbach). Brazil: deep pit galls on the leaves, Rio Grande do Sul, Caxias do Sul, Ana Rech, Faxinal, 700 m, 9.xi.1988, L ithrea brasiliensis (R. Wasum et al.) ; pit galls on the leaves, Bele Âm Ve Âlho, Arraial da Gloria, 31.x.1897, L ithrea molleoides . Paraguay: deep pit galls on the leaves and larvae, Canendiyu, Sierra de Marayu Â, road to Yerbales, near San Estanislao, 24ss39¾S 56ss26¾W, L ithrea molleoides (E. Hassler) ; deep pit galls on the leaves, Caaguazu Â, San Jose Â, 25ss33¾S 56ss45¾W, in the ®elds, 9.xi.1882, L ithrea molleoides (B. Balansa) . Uruguay: pit galls on the leaves, Lavalleja, Minas, 34ss23¾S 55ss14 ¾W, Penitente, 200 m, 1± 2.xi.1931, L ithrea molleoides ; pit galls on the leaves, Rivera, 30ss54 ¾S 55ss31 ¾W, Galgo, 250± 300 m, 2± 4.xii.1933, L ythrea molleoides (W. G. Herter) ; pit galls on the leaves, Tacuarembo Â, 31ss44¾S 55ss59¾W, x.1895, L ithrea molleoides (Arechavaleta) . Preserved dry and in alcohol (CJBG, NHMB, SABC).