Lepanus villosus Matthews, 1974,

Gunter, Nicole L. & Weir, Thomas A., 2019, Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups, Zootaxa 4695 (3), pp. 201-252: 239-247

publication ID

https://doi.org/10.11646/zootaxa.4695.3.1

publication LSID

lsid:zoobank.org:pub:32E17629-F29A-47C8-ACCB-69A32C17B147

persistent identifier

http://treatment.plazi.org/id/03ED0167-FFA3-FF93-FF1D-F790FBA1FD67

treatment provided by

Plazi

scientific name

Lepanus villosus Matthews, 1974
status

 

Lepanus villosus Matthews, 1974 

( Figs 16A, 16CView FIGURE 16, 17DView FIGURE 17, 18DView FIGURE 18, 19DView FIGURE 19)

Lepanus villosus Matthews, 1974: 118  .

Types examined: Holotype: ♂ “ Palmerston N.P., Q. 1.iv.68, E. Matthews ” / “ Lepanus villosus Det. 1972  M, E.G. Matthews, HOLOTYPE ” / “ ANIC Database No. 25 057986” in ANIC [17°34’0”S, 145°45’0”E]GoogleMaps  . Paratypes: “Palmerston N.P., Q. 1.iv.68, E. Matthews ” / “ Lepanus villosus Det. 1972  M, E.G. Matthews, PARATYPE” / “ ANIC Database No. 25 057987” 2 specimens ♀ in ANIC [17°34’0”S, 145°45’0”E],GoogleMaps  ; “Tully, N.Q., 12.x.1969, R.J. Huppatz ” / “ Lepanus villosus Det. 1972  M, E.G. Matthews, PARATYPE” / “ ANIC Database No. 25 057988” 1 specimen ♀ in ANIC [17°36’0”S, 145°55’0”E]GoogleMaps  ; “QLD. <50m Seymour Ra., 10/vii1971, Taylor Feehan” / Berlesate anic 376, rainforest 17.26x146.0” / “ Lepanus villosus Det. 1972  , E.G. Matthews, PARATYPE” / “ ANIC Database No. 25 057989” 1 specimen ♂ in ANIC [17°26’0”S, 146°0’0”E]GoogleMaps  ; “ Gillies H’way, N.Q. 2/10. GB.” / “ PARATYPE ” / “EM.117.” / “ Lepanus villosus Det. 1972  , E.G. Matthews, PARATYPE” / “ ANIC Database No. 25 057990” 1 specimen ♂ in ANIC [17°12’0”S, 145°42’0”E]GoogleMaps  ; “ QLD. 750m Koombooloomba , 4/vii/1971 Taylor Feehan ” / “Ber- lesate ANIC 259View Materials rainforest” / “ Lepanus villosus Det. 1972  , E.G. Matthews, PARATYPE” / “ ANIC Database No. 25 057991” 1 specimen ♂ in ANIC [17°49’0”S, 145°33’0”E]GoogleMaps  ; Davies Creek , Atherton Tbld, N.Qld, 10 vii 1970, J.H. Connell ” / “ Lepanus villosus Det. 1972  M, E.G. Matthews, PARATYPE” (25-058496 1 specimen ♀, 25-058497 1 specimen ♂, 25-058498 1 specimen ♂, 1 specimen ♀ (on same card), 25-058499 1 specimen ♀, 25-058500 1 speci- men ♂ in QM) [16°59’0”S, 145°33’0”E]GoogleMaps  .

Other material examined. 532 specimens from 81 localities were examined. Cape Tribulation transect Site 5A, 16°4’41”S, 145°27’0”E, 1–31.i.1983, ( T44339View Materials 1 specimen in QM)GoogleMaps  ; Cape Tribulation transect Site 5A, 16°4’41”S, 145°27’0”E, 5–9.i.1983, ( T44480View Materials T44482View Materials 3 specimens in QM)GoogleMaps  ; Cape Tribulation transect Site 5A, 16°4’41”S, 145°27’0”E, 20–23.iv.1983, ( T44483View Materials T44484View Materials 2 specimens in QM)GoogleMaps  ; 1.5 km Eby, N Mount Sorrow , [16°5’0”S, 145°27’0”E], 25.iii.1984, A. Calder & T.A. Weir (25-059588 2 specimens in ANIC)GoogleMaps  ; Thompson Creek, ( Grove ), 16°6’31”S, 145°26’25”E, 12.ii–15.iii.1998, ( T85668View Materials 1 specimen in QM)GoogleMaps  ; Thompson Creek, ( Grove ), 16°6’31”S, 145°26’25”E, 15.iii–7.v.1998, ( T88828View Materials 1 specimen in QM)GoogleMaps  ; Cooper Creek, ( Grove ), 16°9’10”S, 145°24’19”E, 11.i–10.ii.1998, ( T73113View Materials T73119View Materials 7 specimens in QM)GoogleMaps  ; Cooper Creek, ( Grove ), 16°9’10”S, 145°24’19”E, 10.ii– 16.iii.1998, ( T85663View Materials T85667, T85669, T85670, T85672 T85673, T88823 T88825View Materials 12 specimens in QM)GoogleMaps  ; Cooper Creek, ( Grove ), 16°9’10”S, 145°24’19”E, 16.iii –7.v.1998, ( T88819View Materials T88821View Materials 3 specimens in QM)GoogleMaps  ; Cooper Creek, ( Grove ), 16°9’10”S, 145°24’19”E, 11.ix–24.x.1998, ( T88822, T88826 2 specimens in QM)GoogleMaps  ; Pimm’s Block, ( Grove ), 16°11’20”S, 145°24’18”E, 13.iii–8.v.1998, ( T88829, T88831 T88833View Materials 4 specimens in QM)GoogleMaps  ; Pimm’s Block, ( Grove ), 16°11’20”S, 145°24’18”E, 10.ix–24.x.1998, ( T88830, T88834 2 specimens in QM)GoogleMaps  ; Hutchinson Creek, ( Grove ), 16°13’4”S, 145°23’57”E, 13.i–11.ii.1998, ( T73120View Materials T73121View Materials 2 specimens in QM)GoogleMaps  ; Hutchinson Creek, ( Grove ), 16°13’4”S, 145°23’57”E, 11.ii–14.iii.1998, ( T85671View Materials 1 specimen in QM)GoogleMaps  ; Hutchinson Creek, ( Grove ), 16°13’4”S, 145°23’57”E, 14.iii–8.v.1998, ( T88827View Materials 1 specimen in QM)GoogleMaps  ; Alexandra Bay , [16°12’0”S, 145°28’0”E], 24.vi.1971, R.W. Taylor & J. Feehan, (25-057996 1 specimen in ANIC)GoogleMaps  ; Lync-Haven, Daintree area , 16°12’7”S, 145°24’32”E, 35 m, 2–10.xii.2012, T.A. Weir & G.B. Monteith, (25-057655 1 specimen in ANIC)GoogleMaps  ; Lync-Haven, Daintree area , 16°12’7”S, 145°24’32”E, 35 m, 6–9.xii.2012, G.B. Monteith (25-057653 1 specimen in ANIC)GoogleMaps  ; Thornton Range near Daintree , [16°14’0”S, 145°24’0”E], 200 m, 12–18.vii.1982, S. & J. Peck (25-059578 6 specimens in ANIC)GoogleMaps  ; Buchanan Creek, ( Grove ), 16°14’39”S, 145°25’54”E, 12.iii–8.v.1998, ( T88818View Materials 1 specimen in QM)GoogleMaps  ; Humbug Landing, Daintree , 16°15’13”S, 145°20’54”E, 14–18.v.1998, ( T66779View Materials T66780View Materials 2 specimens in QM)GoogleMaps  ; Mount Gorton , 16°57’0”S, 145°53’0”E, 18.xi.1993, ( T44315View Materials T44321View Materials 7 specimens in QM)GoogleMaps  ; Mount Murray Prior , 16°56’0”S, 145°51’0”E, 7.xii.1995 ( T44324View Materials 1 specimen in QM)GoogleMaps  ; Mount Murray Prior , 16°56’0”S, 145°51’0”E, 31.x.1995 – 8.xii.1995, ( T44419View Materials T44423View Materials 5 specimens in QM)GoogleMaps  ; Mount Murray Prior , 16°56’0”S, 145°51’0”E, 7–8.xii.1998, ( T70933View Materials T70954View Materials 21 specimens in QM)GoogleMaps  ; Chujeba Peak summit, 16°56’14”S, 145°39’19”E, 16.xii.1989, ( T44340View Materials 1 specimen in QM)GoogleMaps  ; Davies Creek Road via Mareeba , [17°1’0”S, 145°35’0”E], 9.x–17.x.1976, R.I. Storey (25-059595 3 specimens in ANIC, 2 specimens in QDPC)GoogleMaps  ; Davies Creek Road, via Mareeba , 17°2’33”S, 145°36’51”E, 26.xi.1985, ( T44341View Materials x 1 in QM)GoogleMaps  ; Davies Creek Road, via Mareeba , 17°2’33”S, 145°36’51”E, 4–13.xii.1988, ( T44414View Materials T44415View Materials 2 specimens in QM)GoogleMaps  ; Isley Creek , upper, 17°2’41”S, 145°39’57”E, 29–30.xi.1993, ( T44330View Materials T44338View Materials 9 specimens in QM)GoogleMaps  ; Mount Tiptree , [17°3’0”S, 145°37’0”E], 13.vii.1984, R.B. Halliday (25-059585 1 specimen in ANIC)GoogleMaps  ; Mount Tiptree , 17°3’0”S, 145°38’0”E], 730 m, 29.vi.1971, R.W. Taylor & J. Feehan SAMA25- 027824GoogleMaps  2 specimens in SAMA  ); North Bell Peak summit, 17°5’19”S, 145°52’44”E, 16.ix.1981, ( T44328View Materials 1 specimen in QM)GoogleMaps  ; North Bell Peak summit, 17°5’19”S, 145°52’44”E, 22.xi.1990, ( T44342View Materials T44345View Materials 3 specimens in QM)GoogleMaps  ; North Bell Peak summit, 17°5’19”S, 145°52’44”E, 20–22.xi.1990, ( T44365View Materials T44370View Materials 6 specimens in QM)GoogleMaps  ; Lamb Range , 6 km NNE Tinaroo Falls, 17°6’40”S, 145°33’58”E, 1159 m, 16.xi.2009, G.B. Monteith & Turco, (25- 060851 3 specimens in ANIC aslso as COL960- COL962)GoogleMaps  ; Danbulla State Forest Robson Creek Road 1.5 km N of road to Tinaroo , [17°7’0”S, 145°37’0”E], 27.iv–2.v.1998, Miledge,G., (25- 060767 1 specimen in AM)GoogleMaps  ; Danbulla S.F., 13 km NE of Yungaburra [Danbulla Forestry Reserve], [17°10’0”S, 145°38’0”E], 13.i–13.ii.1987, R.I. Storey & De Faveri, (25-059594 1 specimen in QDPC)GoogleMaps  ; Danbulla State Forest , 13 km NE of Yungaburra [Danbulla Forestry Reserve], [17°10’0”S, 145°38’0”E], 23.iv–27.v.1987, R.I. Storey & De Faveri, (25-059604 1 specimen in ANIC)GoogleMaps  ; Danbulla State Forest , 13 km NE of Yungaburra [Danbulla Forestry Reserve], [17°10’0”S, 145°38’0”E], 23.vi–28.vii.1987, R.I. Storey & De Faveri, (25- 032825 1 specimen in QDPC)GoogleMaps  ; Danbulla State Forest , 13 km NE of Yungaburra [Danbulla Forestry Reserve], [17°10’0”S, 145°38’0”E], 28.vii–3.ix.1987, R.I. Storey & De Faveri, (25- 032826 1 specimen in QDPC)GoogleMaps  ; Cathedral Fig , 17°10’52”S, 145°39’26”E, 10.xii.1988, ( T44326View Materials 1 specimen in QM)GoogleMaps  ; Graham Range , summit, 17°16’24”S, 145°57’58”E, 1.xi–8.xii.1995, ( T44404View Materials T44405View Materials 2 specimens in QM)GoogleMaps  ; Graham Range , 350 m, 17°16’37”S, 145°57’48”E, 1.xi–8.xii.1995, ( T54656View Materials T54657View Materials 2 specimens in QM)GoogleMaps  ; Boar Pock- et Road , approximately 8 km N of Gillies Highway [Gillies Road], [17°12’0”S, 145°42’0”E], 13.xii.1969, J.G. Brooks, (25-057998 1 specimen in ANIC)GoogleMaps  ; Boar Pocket Road , approximately 8 km N of Gillies Highway [Gillies Road], [17°12’0”S, 145°42’0”E], 9.xii.1969, J.G. Brooks, (25-057997 1 specimen in ANIC)GoogleMaps  ; Boar Pocket Road 3–9 mi, 17°11’57”S, 145°40’8”E, 2500 feet, 8.iv.1974, J.G. Brooks, (CMNEN-00030586-00030588 in CMNC)GoogleMaps  ; Bellenden Ker Range, 0.5 km S Cable Tower 7, [17°16’0”S, 145°55’0”E], 500 m, 25–31.x.1981, Earthwatch, Queensland Museum , (25- 032830 1 specimen in QDPC, 25-059582 x 5 in ANIC)GoogleMaps  ; Bellenden Ker Range , 1 km S Cable Tower 6, 17°16’0”S, 145°53’0”E], 500 m, 17–24.x.1981, Earthwatch, Queensland Museum, (25-057999 2 specimens in ANIC)GoogleMaps  ; Bellenden Ker Range , 1 km S Cable Tower 6, 17°16’0”S, 145°53’0”E], 500 m, 25–31.x.1981, Earthwatch, Queensland Museum, (25-059583 1 specimen in ANIC)GoogleMaps  ; Lake Barrine National Park , [17°16’0”S, 145°39’0”E], 7.vii.1984, R.B. Halliday (25-059587 1 specimen in ANIC)GoogleMaps  ; Wright Creek, Lake Eacham , [17°16’0”S, 145°37’0”E], 750 m, 31.vii.1982, S. & J. Peck (25-058000 2 specimens in ANIC)GoogleMaps  ; Wright Creek, Lake Eacham , [17°16’0”S, 145°37’0”E], 720 m, 26–30.vii.1982, S. & J. Peck (25-059579 2 specimens in ANIC)GoogleMaps  ; Baldy Mountain Road Atherton , [17°17’0”S, 145°24’0”E], 12.xii.1976, R.I. Storey (25- 032828 1 specimen in QDPC)GoogleMaps  ; Baldy Mountain near Atherton [Mount Baldy], [17°17’0”S, 145°27’0”E], xii.1977 – i.1978, R.I. Storey (25- 032827 2 specimens in QDPC)GoogleMaps  ; Eacham National Park [Lake Eacham National Park], [17°17’0”S, 145°37’0”E], 760 m, 21.iii.1975, R.W.Taylor (25-057994 1 specimen in ANIC)GoogleMaps  ; Bartle Frere, Gourka Road , 17°22’38”S, 145°44’34”E, 28–30.xi.1998, ( T70929View Materials T70932View Materials 4 specimens in QM)GoogleMaps  ; Bartle Frere, West Base , 17°22’47”S, 145°45’51”E, 10.i–31.iii.1995, ( T44447View Materials T44452View Materials 6 specimens in QM)GoogleMaps  ; Bartle Frere, West Base , 17°22’47”S, 145°45’51”E, 7.iii–15.v.1995, ( T44453View Materials T44464, T55904 11 specimens in QM)GoogleMaps  ; Bartle Frere, West Base , 17°22’47”S, 145°45’51”E, 25.xi.1994 – 10.i.1995, ( T44465View Materials T44479View Materials 15 specimens in QM)GoogleMaps  ; Lamins Hill , 17°22’56”S, 145°42’7”E, 1.xii.1993 – 25.ii.1994, ( T44413View Materials 1 specimen in QM)GoogleMaps  ; Bartle Frere W Base , [17°23’0”S, 145°46’0”E], 750 m, 28.xi.1998, G.B. Monteith (25- 060849 1 specimen in ANIC)GoogleMaps  ; Mount Bartle Frere W side [Bartle Frere South Peak], 17°24’0”S, 145°49’0”E], 700 m, 30.vii.1982, S. & J. Peck (25-059580 2 specimens in ANIC)GoogleMaps  ; Mount Bartle Frere W side [Bartle Frere South Peak], [17°24’0”S, 145°49’0”E], 700 m, 30.vii–4.viii.1982, S. & J. Peck (25-059581 1 specimen in ANIC)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 12.x–5.xi.1983, R.I. Storey & Brown (25-059601 1 specimen in ANIC)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 13.iii–1.v.1984, R.I. Storey & Brown (25-059603 1 specimen in QDPC)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 21.vi–26.vii.1984, R.I. Storey & Brown (25-059602 x 1 in ANIC)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 5.xi–1.xii.1983, R.I. Storey & Brown (25- 032829 1 specimen in QDPC)GoogleMaps  ; Topaz, Hughes Road, ( Breeden house), 17°26’4”S, 145°42’2”E, 1.vii–30.ix.1993, ( T44426View Materials 1 specimen in QM)GoogleMaps  ; 21 km S of Atherton , 17°26’41”S, 145°28’35”E, 5–14.xii.1988, ( T44416View Materials 1 specimen in QM)GoogleMaps  ; Hugh Nelson Range , [17°27’0”S, 145°29’0”E], 1150 m, 3.vii–1.viii.1995, P. Zborowski (25-058038 1 specimen in ANIC)GoogleMaps  ; Josephine Falls, Mount Bartle Frere [Bartle Frere], [17°27’0”S, 145°52’0”E], 10.vii.1984, R.B. Halliday (25-059586 2 specimens in ANIC)GoogleMaps  ; Longlands Gap , 17°27’14”S, 145°28’26”E, 1120 m, 18–19.ix.2010, G.B. Monteith (25-059606 1 specimen, 25- 060866 3 specimens in ANIC also as COL1123-COL1125)GoogleMaps  ; tower near The Crater , 17°27’22”S, 145°29’13”E, 1235m, 21–23.i.2012, G.B. Monteith (25-055609 7 specimens in ANIC, 25-053847 2 specimens in ANIC)GoogleMaps  ; Tower S of Crater National Park, 17°27’23”S, 145°29’12”E, 25.xi.1994 – 10.i.1995, ( T44346View Materials T44351View Materials 6 specimens in QM)GoogleMaps  ; Tower S of Crater National Park, 17°27’23”S, 145°29’12”E, 10.i–31.iii.1995, ( T44352View Materials T44359View Materials 8 specimens in QM)GoogleMaps  ; Tower S of Crater National Park, 17°27’23”S, 145°29’12”E, 7.iii–15.v.1995, ( T44360View Materials T44364View Materials 5 specimens in QM)GoogleMaps  ; Longlands Gap , 17°27’46”S, 145°29’32”E, 30.x.1991 – 23.vii.1992, ( T44389View Materials 1 specimen in QM)GoogleMaps  ; Millaa Millaa Falls (below), [17°29’0”S, 145°36’0”E], 760 m, 18–20.iv.1999, C. & I. Reid (25-059605 1 specimen in ANIC)GoogleMaps  ; Millaa Millaa Falls , 17°29’53”S, 145°36’32”E, 6–9.ii.1999, ( T76229,T76231 T76232View Materials 3 specimens in QM)GoogleMaps  ; Kjellberg Road turnoff, 17°31’57”S, 145°36’27”E, 7.iii–15.v.1995, ( T44380View Materials 1 specimen in QM)GoogleMaps  ; Mount Fisher, Kjellberg Road , 17°32’34”S, 145°33’31”E, 27.iv.1982, ( T44322View Materials T44323View Materials 2 specimens in QM)GoogleMaps  ; Mount Fisher, Kjellberg Road , 17°32’34”S, 145°33’31”E, 3.v.1983, ( T44329View Materials 1 specimen in QM)GoogleMaps  ; Mount Fisher, Kjellberg Road , 17°32’34”S, 145°33’31”E, 27–29.iv.1982, ( T44417View Materials T44418, T44425 3 specimens in QM)GoogleMaps  ; Kjellberg Road, State Forest boundary, 17°32’9”S, 145°34’39”E, 6–9.ii.1999, ( T76233View Materials 1 specimen in QM)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 10.i–31.iii.1995, ( T44427View Materials T44431View Materials 5 specimens in QM)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 7.iii–15.v.1995, ( T44435View Materials T44438View Materials 4 specimens in QM)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 25.xi.1994 – 10.i.1995, ( T44439View Materials T44446View Materials 8 specimens in QM)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 6–9.ii.1999, ( T76219View Materials T76227View Materials 9 specimens in QM)GoogleMaps  ; Henrietta Creek Palmerston National Park , [17°35’54”S, 145°45’30”E], 22–24.iv.1973, R.I. Storey (CMNEN-00030582-00030585 in CMNC)GoogleMaps  ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 11.v.1974, R.I. Storey (25- 032831 1 specimen in QDPC)GoogleMaps  ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 13. vii–15.viii.1990, R.I. Storey & H.K. Halfpapp, (25- 032818 3 specimens in QDPC)GoogleMaps  ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 13.iii–10.iv.1990, R.I. Storey & H.K. Halfpapp, (25- 032816 3 specimens in QDPC)GoogleMaps  ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 15.viii–12.ix.1990, R.I. Storey & H.K. Halfpapp, (25- 032819 2 specimens in QDPC)GoogleMaps  ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 4.i–7.ii.1990, R.I. Storey & H.K. Halfpapp, (25- 032815 1 specimen in QDPC)GoogleMaps  ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 6.vi–13.vii.1990, R.I. Storey & H.K. Halfpapp, (25- 032817 2 specimens in QDPC)GoogleMaps  ; BS2View Materials Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 31.i–27.ii.1996, L. Umback (25-059610 1 specimen in ANIC)GoogleMaps  ; Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 1.xii.1994 – 3.i.1995, P. Zborowski (25-059613 1 specimen in ANIC)GoogleMaps  ; Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 2.viii–4.ix.1995, L. Umback (25-058007 3 specimens in ANIC)GoogleMaps  ; Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 2.vi–4.vii.1995, P. Zborowski (25-058036 1 specimen in ANIC)GoogleMaps  ; Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 2–30.xi.1995, L. Umback (25-058013 1 specimen in ANIC)GoogleMaps  ; Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 21.iii–5.iv.1995, P. Zborowski (25-058021 1 specimen, 25-058022 2 specimens, 25-058023 2 specimens, 25-058024 3 specimens in ANIC)GoogleMaps  ; Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 3.i–4.ii.1995, P. Zborowski (25-059611 2 specimens in ANIC)GoogleMaps  ; Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 3.v–2.vi.1995, P. Zborowski (25-058034 3 specimens, 25-058035 4 specimens in ANIC)GoogleMaps  ; Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 3.x–2.xi.1995, L. Umback (25-058011 2 specimens in ANIC)GoogleMaps  ; Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 4.ii–21.iii.1995, P. Zborowski (25-059612 2 specimens in ANIC)GoogleMaps  ; Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 4.vii–2.viii.1995, P. Zborowski (25-058001 2 specimens, 25-058002 1 specimen in ANIC)GoogleMaps  ; Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 4.ix–3.x.1995, L. Umback (25-058008 1 specimen, 25-058009 9 specimens in ANIC)GoogleMaps  ; Mount Fisher , [17°33’0”S, 145°32’0”E], 1150 m, 5.iv–3.v.1995, P. Zborowski (25-058025 3 specimens, 25-058026 5 specimens, 25-058027 3 specimens, 25-058028 1 specimen, 25-058029 3 specimens, 25-058030 2 specimens in ANIC)GoogleMaps  ; Henrietta Creek Palmerston National Park [Plamerston National Park], [17°34’0”S, 145°45’0”E], 22–24.iv.1973, R.I. Storey (25- 032824 6 specimens in QDPC, 25-058501 8 specimens in QM)GoogleMaps  ; BS3 Massey Creek , [17°37’0”S, 145°34’0”E], 1000 m, 1.xii.1994 – 3.i.1995, P. Zborowski (25-059608 5 specimens in ANIC)GoogleMaps  ; BS3View Materials Massey Creek , [17°37’0”S, 145°34’0”E], 1000 m, 3.i–4.ii.1995, P. Zborowski (25-059609 1 specimen in ANIC)GoogleMaps  ; BS3View Materials Massey Creek , [17°37’0”S, 145°34’0”E], 1000 m, 4.ii–6.iii.1995, P. Zborowski (25-059614 3 spec- imens in ANIC)GoogleMaps  ; Massey Creek , [17°37’0”S, 145°34’0”E], 1000 m, 2.viii–4.ix.1995, L. Umback (25-058006 6 specimens in ANIC)GoogleMaps  ; Massey Creek , [17°37’0”S, 145°34’0”E], 1000 m, 2.vi–4.vii.1995, P. Zborowski (25-058037 1 specimen, 25-058039 5 specimens, 25-058040 5 specimens in ANIC)GoogleMaps  ; Massey Creek , [17°37’0”S, 145°34’0”E], 1000 m, 2.xi–30.xi.1995, L. Umback (25-058014 1 specimen, 25-058015 3 specimens in ANIC)GoogleMaps  ; Massey Creek , [17°37’0”S, 145°34’0”E], 1000 m, 3.v–2.vi.1995, P. Zborowski (25-058031 1 specimen, 25-058032 2 specimens, 25-058033 2 specimens in ANIC)GoogleMaps  ; Massey Creek , [17°37’0”S, 145°34’0”E], 1000 m, 3.x–2.xi.1995, L. Umback (25-058012 14 specimens in ANIC)GoogleMaps  ; Massey Creek , [17°37’0”S, 145°34’0”E], 1000 m, 4.vii–2.viii.1995, P. Zborowski (25-058003 6 specimens, 25-058004 3 specimens, 25-058005 2 specimens in ANIC)GoogleMaps  ; Massey Creek , [17°37’0”S, 145°34’0”E], 1000 m, 4.ix–3.x.1995, L. Umback (25-058010 9 specimens in ANIC)GoogleMaps  ; Massey Creek , [17°37’0”S, 145°34’0”E], 1000 m, 6.iii–5.iv.1995, P. Zborowski (25-058016 3 specimens, 25-058017 1 specimen, 25-058018 2 specimens, 25-058019 12 specimens, 25-058020 17 specimen in ANIC)GoogleMaps  ; South Johnstone Forestry Camp , 5 km W, 17°38’12”S, 145°40’15”E, 1.xii.1993 – 25.ii.1994, ( T44403View Materials 1 specimen in QM)GoogleMaps  ; Vine Creek, Majors Mountain , 17°40’58”S, 145°32’2”E, 4–6.ii.1999, ( T76234View Materials T76236View Materials 3 specimens in QM)GoogleMaps  ; Vine Creek, Majors Mountain , 17°40’58”S, 145°32’2”E, 15–18.iv.1999, ( T79131View Materials 1 specimen in QM)GoogleMaps  ; Charmillin Creek crossing, 17°42’3”S, 145°31’21”E, 8.xii.1989 – 5.i.1990, ( T44373View Materials T44374View Materials 23 specimens in QM)GoogleMaps  ; Charmillin Creek crossing, 17°42’3”S, 145°31’21”E, 5–11.ii.1998, ( T62197View Materials 1 specimen in QM)GoogleMaps  ; Ravenshoe State Forest, Tully Falls Road, [17°43’0”S, 145°33’0”E], 10.x–15.xi.1987, A. Walford-Huggins (25-059584 2 specimens in ANIC)GoogleMaps  ; Ravenshoe State Forest, Tully Falls Road, [17°43’0”S, 145°33’0”E], 15.xi–12.xii.1987, A. Walford-Huggins (25-059589 3 specimens in ANIC)GoogleMaps  ; Tully Falls State Forest 18 km SSW Ravenshoe, [17°46’0”S, 145°31’0”E], 730 m, 1.x– 5.xi.1987, R.I. Storey & Dickinson, (25- 032820 1 specimen in QDPC, 25-059590 6 specimens in ANIC)GoogleMaps  ; Tully Falls State Forest 18 km SSW Ravenshoe, [17°46’0”S, 145°31’0”E], 730 m, 20.iv–25.v.1988, R.I. Storey & Dickinson, (25-059591 1 specimen in ANIC)GoogleMaps  ; Tully Falls State Forest 18 km SSW Ravenshoe, [17°46’0”S, 145°31’0”E], 730 m, 5.i–7.ii.1988, R.I. Storey & Dickinson, (25- 032822 1 specimen in QDPC, 25-059598 1 specimen in ANIC)GoogleMaps  ; Tully Falls State Forest 18 km SSW Ravenshoe, [17°46’0”S, 145°31’0”E], 730 m, 7.xii.1987 – 7.i.1988, R.I. Storey & Dickinson, (25-059597 2 specimens in ANIC)GoogleMaps  ; Tully Falls State Forest 18 km SSW Ravenshoe, [17°46’0”S, 145°31’0”E], 730 m, 7.xi–7.xii.1987, R.I. Storey & Dickinson, (25- 032821 2 specimens in QDPC, 25-059592 4 specimens in ANIC)GoogleMaps  ; Tully Falls State Forest 18 km SSW Ravenshoe, [17°46’0”S, 145°31’0”E], 730 m, 9.ii– 3.iii.1988, R.I. Storey & Dickinson, (25-059600 2 specimens in ANIC)GoogleMaps  ; Tully Falls State Forest 11 km SSW Ravenshoe, [17°47’0”S, 145°34’0”E], 900 m, 1.x–5.xi.1987, R.I. Storey & Dickinson, (25- 032823 1 specimen in QDPC, 25-059593 1 specimen in ANIC)GoogleMaps  ; Tully Falls State Forest 11 km SSW Ravenshoe, [17°47’0”S, 145°34’0”E], 900 m, 20.iv–25.v.1988, R.I. Storey & Dickinson, (25-059599 1 specimen in ANIC)GoogleMaps  ; Tully Falls State Forest 11 km SSW Ravenshoe, [17°47’0”S, 145°34’0”E], 900 m, 7.xii.1987 – 7.i.1988, R.I. Storey & Dickinson, (25-059596 2 specimens in ANIC)GoogleMaps  ; Tully Falls State Forest via Ravenshoe , [17°47’0”S, 145°34’0”E], 20–21.iv.1973, R.I. Storey (25-058502 1 specimen in QM)GoogleMaps  ; Tully Falls , 17°47’0”S, 145°33’27”E, 8.xii.1989 - 5.i.1990, ( T44379View Materials 1 specimen in QM)GoogleMaps  ; Red Road turnoff on Tully Falls Road , 17°49’19”S, 145°32’42”E, 8.xii. 198 9–5.i.1990, ( T44371View Materials 1 specimen in QM)GoogleMaps  ; Boulder Creek track, 200–600 m, 17°51’2”S, 145°54’29”E, 24–27.x.1983, ( T44390View Materials - T44397View Materials 8 specimens in QM)GoogleMaps  ; Laceys Creek , [17°52’0”S, 146°4’0”E], 40 m, 11.vii.1971, R.W. Taylor J.E. Feehan, (25-057995 1 specimen in ANIC)GoogleMaps  ; Tully , [17°53’0”S, 145°55’0”E], 200 m, 17.x–22.xi.1994, (25-059607 1 specimen in ANIC)GoogleMaps  ; Tully River Crossing , 10 km S Koombooloomba Dam, 17°55’32”S, 145°37’38”E, 8.xii.1989 – 4.i.1990, ( T44375View Materials T44378View Materials 4 specimens in QM)GoogleMaps  ; Mount Tyson , 17°55’35”S, 145°54’34”E, 7.v.1983, ( T44327View Materials 1 specimen in QM)GoogleMaps 

Description. Uniformly brown in colour. Antennal clubs yellow to white.

Total length: 2.2–2.8 mm. Holotype measurements: total length 2.4 mm, elytral width 1.5 mm.

Male. Head. Surface smooth and nitid, with dense ocellate punctures each bearing a seta. Very broad U-shaped between clypeal teeth that are short but not upturned. Genal angle weakly defined. Dorsal part of the eyes narrow, separated by an interocular space approximately 11 times the eye width (34:3). Head 1.5 x wider than long (45:30).

Prothorax: Pronotal surface smooth and nitid, with dense ocellate punctures each bearing an obvious posteiorly directed seta. Punctures along the basal margin horseshoe shaped differing from those on disc. Anterior angles obtuse. Basal margin slightly angulate. Hypomeral surface reticulate and punctate. Hypomeral stria about two thirds the lenth of hypomeron with a row of fine setae present between hypomeral stria and lateral edge. Pronotum 1.8

x wider than long (68:38).

Elytra: Intervals reticulate with fine meshes, and with a double row of recurved setae. Striae geminate, appearing double. A well-developed humeral callus between striae 6 and 7, and a basal depression between striae 4 and 5. Stria 8 ending basally in a deep pit and lateral carina indented at this point. Elytra 1.05 x wider than long (87:82).

Legs: Apical digit of foretibiae broad with a small comb of setae on inner apex and very small apical spur on outer edge. Metatibiae with a notch on the inner edge.

Abdomen: Pygidium flat, reticulate, punctate and with setae and with an overhanging basal edge. Abdominal ventrites reticulate right across.

Pterothorax: Medial lobe of metaventrite punctate and narrowly margined between the mesocoxae. Lateral lobe of metaventrite with ocellate punctures. Meso-metaventral suture very slightly arched. Mesoventrite smooth with punctation. Mesepimeron reticulate and punctate. Metanepisternum reticulate and punctate.

Hind wings: Fully developed.

Female. Metatibae without a notch on the inner edge, otherwise as per the L. villosus  species group.

Distribution. Known from rainforests of the Australian Wet Tropics bioregion from Cape Tribulation to Tully ( Fig. 20View FIGURE 20). The majority of specimens have been collected from mid to high elevation forest (500–1200 m) but some are known from forest at sea level.

Comments. Almost 550 specimens were examined for this revision of which almost all have been collected in flight intercept traps, pitfall traps, or in berlesed leaf and log litter. The few specimens that have data for baited pitfalls indicate a preference of kangaroo dung, but other attractants include human faeces, carrion, mushroom, and fish baits. Ebert et al. (2019) only collected a single specimen of L. villosus  in their 10-way food preference study and it came to a kangaroo dung baited pitfall trap. Specimens collected from Lambs Range and Longlands Gap (25- 060866 and 25- 060851) have been sequenced and included in the phylogenetic analyses ( Gunter & Weir 2017; Gunter et al. 2019). In the latter more comprehensively sampled study, L. villosus  is recovered as sister to L. pseudovillosus  new species, in a monophyletic clade that also contained L. vestitus  .

ANIC

Australian National Insect Collection

QM

Queensland Museum

QDPC

Queensland Primary Industries Insect Collection

SAMA

South Australia Museum

AM

Australian Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Lepanus

Loc

Lepanus villosus Matthews, 1974

Gunter, Nicole L. & Weir, Thomas A. 2019
2019
Loc

Lepanus villosus

Matthews, E. G. 1974: 118
1974