Lepanus reidi Gunter & Weir,

Gunter, Nicole L. & Weir, Thomas A., 2019, Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups, Zootaxa 4695 (3), pp. 201-252: 231-234

publication ID

https://doi.org/10.11646/zootaxa.4695.3.1

publication LSID

lsid:zoobank.org:pub:32E17629-F29A-47C8-ACCB-69A32C17B147

persistent identifier

http://treatment.plazi.org/id/03ED0167-FF9B-FF9C-FF1D-F0ACFE19F917

treatment provided by

Plazi

scientific name

Lepanus reidi Gunter & Weir
status

new species

Lepanus reidi Gunter & Weir  , new species

( Figs 8B, F, HView FIGURE 8; 9CView FIGURE 9; 10CView FIGURE 10; 11CView FIGURE 11.)

Type series. Holotype: ♂ “QLD: 17.430°S 145.490°E Crater NP, 18-19 Sep 2010, G. Monteith Dung trap, RF, 980m 34584” (= Mout Hypipamee National Park ) / “ ANIC Database No. 25 059231” / “ ANIC image” / “ Lepanus  NQ5 det TA Weir 2010 ”, in ANIC (17°25’47”S, 145°29’24”E)GoogleMaps  . Paratypes (25 ♂, 35 ♀, 5 sex undetermined): Crater National Park , 980 m, dung trap 34584, 17.430°S, 145.490°E 18–19.ix.2010, G.B. Monteith (25- 060883 3 specimens in ANIC also as COL1152 ♀, COL1153 ♀, COL1154 ♀) (17°25’47”S, 145°29’24”E)GoogleMaps  ; Mount Hartley , 2.5 km SW, 15°47’49”S, 145°19’34”E, 24.iv.1982, ( T 54260View Materials in QM 1 specimen ♀)GoogleMaps  ; Mount Finnigan S1, 30 km S of Cooktown, [15°49’0”S, 145°17’0”E], 400 m, 2.vii.1982, S. & J. Peck (25-059106 1 specimen ♂ 1 specimen ♀ in ANIC)GoogleMaps  ; Alexandra Range , 16°14’42”S, 145°26’10”E, 200 m, 20–21.ix.2010, G.B. Monteith (25- 060882 1 specimen in ANIC also as COL1151 ♀)GoogleMaps  ; Mount Sorrow track, 16°4’37”S, 145°27’58”E, 30 m, 28–30.v.2012, G.B. Mon- teith (25-057612 1 specimen, 25-057617 1 specimen ♀ also as COL 2448in ANIC)GoogleMaps  ; Emmagen Creek , 16°2’24”S, 145°27’32”E, 10 m, 29–30.v.2012, G.B. Monteith (25-057618 2 specimens ♀ also as COL 2450 in ANIC)GoogleMaps  ; 1 km WNW Cape Tribulation , [16°4’0”S, 145°28’0”E], 10 m, x.1996, J. Seymour (25-059136 1 specimen ♀ in ANIC)GoogleMaps  ; Cape Tribulation , [16°5’0”S, 145°27’0”E], 20 m, 9.v.1997, C. Reid (25-059134 1 specimen ♀, 25-059135 1 speci- men ♀ in ANIC)GoogleMaps  ; Cape Tribulation , 40 km N of Daintree, [16°5’0”S, 145°29’0”E], 10 m, 12–15.vii.1982, S. & J. Peck (25-059105 3 specimens ♂, 2 specimens ♀ in ANIC)GoogleMaps  ; Oliver Creek , 16°8’22”S, 145°26’24”E, 4–7.vi.1974, ( T60127View Materials 1 specimen ♂ in QM)GoogleMaps  ; Alexandra Range , N slope, 16°14’31”S, 145°25’48”E, 140 m, 28–30.v.2012, G.B. Monteith (25-057616 1 specimen ♂ also as COL 2449 in ANIC)GoogleMaps  ; Windsor Tableland ANZSES Site 9, 16°16’55”S, 145°3’56”E, 27.xii.1988 – 8.i.1989, ( T54257View Materials 1 specimen ♀ in QM)GoogleMaps  ; Mossman Bluff Track 5–10 km W Mossman, Site 1, 16°28’19”S, 145°19’37”E, 20.xii.1989 – 15.i.1990, ( T54269View Materials T54270View Materials 1 specimen ♀, 1 specimen sex undeter- mined in QM)GoogleMaps  ; Mossman Bluff Track 5–10km W Mossman, Site 1, 16°28’19”S, 145°19’37”E, 1.i.1989 – 16.i.1989, ( T54271View Materials T54279View Materials 7 specimens ♂, 2 specimens ♀ in QM)GoogleMaps  ; Mossman Gorge, 16°28’26”S, 145°19’54”E, 1.xi.1991 – 22.vii.1992, ( T54258View Materials T54259View Materials 1 specimens ♂, 1 specimens ♀ in QM)GoogleMaps  ; Black Mountain , lower slopes, 16°39’52”S, 145°30’4”E, 2.xii.1997 – 6.ii.1998, ( T60126View Materials 1 specimen ♀ in QM)GoogleMaps  ; GS1 Mount Haig , [17°6’0”S, 145°36’0”E], 1150 m, 1.xii.1994 – 3.i.1995, P. Zborowski (25-054103 1 specimen ♂, 2 specimen ♀ in ANIC)GoogleMaps  ; Massey Range , 2nd camp, 17°13’51”S, 145°48’13”E, 11–12.x.1991, ( T54256View Materials 1 specimen ♀ in QM)GoogleMaps  ; Bartle Frere , West Base, 17°22’47”S, 145°45’51”E, 25.xi.1994 – 10.i.1995, ( T54261View Materials T 54264View Materials in QM 1 specimen ♂, 3 specimen ♀)GoogleMaps  ; Bartle Frere , West Base, 17°22’47”S, 145°45’51”E, 10.i–31.iii.1995, ( T54265View Materials T54266View Materials 2 specimens ♀ in QM)GoogleMaps  ; Bartle Frere , West Base, 17°22’47”S, 145°45’51”E, 7.iii–15.v.1995, ( T54267View Materials 1 specimen sex undetermined in QM)GoogleMaps  ; BS3 Massey Creek , 17°37’0”S, 145°34’0”E], 1000 m, 1.xii.1994 – 3.i.1995, P. Zborowski (25-054101 3 specimens ♂, 3 speci- mens ♀ in ANIC)GoogleMaps  ; BS3View Materials Massey Creek , 17°37’0”S, 145°34’0”E], 1000 m, 3.x–2.xi.1995, L. Umback (25-054105 1 specimen ♂ in ANIC)GoogleMaps  ; Mount Fisher site BS2, [17°33’0”S, 145°32’0”E], 1150 m, 10.xii.1994 – 3.i.1995, P. Zborows- ki (25-054104 1 specimen ♀ in ANIC)GoogleMaps  ; Maalan Road , 2 km S Palmerston Highway, 17°35’56”S, 145°41’58”E, 7.iii–15.v.1995, ( T54252View Materials T54253View Materials 1 specimen ♂, 1 specimen ♀ in QM)GoogleMaps  ; Maalan Road , 2 km S Palmerston Highway, 17°35’56”S, 145°41’58”E, collected 26.xi.1994 – 10.i.1995, ( T54254View Materials T54255View Materials 1 specimen ♂, 1 specimen sex unde- termined in QM)GoogleMaps  ; Bellenden Ker Range , 0.5 km S Cable Tower 7, [17°16’0”S, 145°55’0”E], 500 m, 25–31.x.1981, Earthwatch, Queesland Museum, (25-059108 1 specimen ♂ in ANIC)GoogleMaps  ; Bellenden Ker Range , Cableway Base Sta- tion, [17°16’0”S, 145°55’0”E], 100 m, 17.x–9.xi.1981, Earthwatch, Queensland Museum, (25-059110 1 specimen sex undetermined in ANIC)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 1.xii.1983 – 9.i.1984, R.I. Storey & Brown (25-059130 1 specimen ♂, 1 specimen ♀ in CMNC)GoogleMaps  ; Mount Father Clancy , 800 m, 17°35’58”S, 145°38’4”E, 6–14.xii.1988, ( T54219View Materials 1 specimen ♀ in QM)GoogleMaps  ; Palmerston National Park , E margin, 17°36’40”S, 145°45’48”E, 2.xi.1995 – 10.xii.1995, ( T54210View Materials 1 specimen ♂ in QM)GoogleMaps  .

Other material examined. 427 specimens from almost 50 localities. Mount Boolbun South , (camp in QM)  , 15°57’10”S, 145°8’’23”E, 5.xi.1995 – 11.i.1996, ( T54191View Materials T 54195View Materials in QM)  ; Mount Boolbun South , (camp in QM)  , 15°57’10”S, 145°8’’23”E, 5.xi.1995 ( T 54196View Materials in QM); Mount Boolbun South , (camp in QM), 15°57’10”S, 145°8’’23”E, 4–6.xi.1995, ( T54197View Materials T 54201View Materials in QM)  ; Donovan Creek, ( Grove ), 16°1’13”S, 145°26’38”E, 14.i– 10.ii.1998, ( T 73143View Materials in QM)GoogleMaps  ; Donovan Creek, ( Grove ), 16°1’13”S, 145°26’38”E, 10.ii.1998 – 14.iii.1998, ( T 85650View Materials in QM)GoogleMaps  ; Donovan Creek, ( Grove ), 16°1’13”S, 145°26’38”E, 14.iii– 6.v.1998, ( T88788View Materials T88790, T88793 T 88794View Materials in QM)GoogleMaps  ; Donovan Creek, ( Grove ), 16°1’13”S, 145°26’38”E, 10.ix–26.x.1998, ( T88791View Materials T 88792View Materials in QM)GoogleMaps  ; Cape Tribulation transect Site 1, 16°4’12”S, 145°27’54”E, 23.ix–7.x.1982, ( T 54165View Materials in QM)GoogleMaps  ; Cape Tribulation transect Site 2, 16°4’28”S, 145°27’33”E, 20–23.iv.1983, ( T54162View Materials T 54164View Materials in QM)GoogleMaps  ; Cape Tribulation transect Site 2, 16°4’28”S, 145°27’33”E, 23.ix –7.x.1982, ( T 54166View Materials in QM)GoogleMaps  ; Cape Tribulation transect Site 5, 16°4’41”S, 145°27’13”E, 20–23.iv.1983, ( T54146View Materials T 54160View Materials in QM)GoogleMaps  ; Cape Tribulation transect Site 5, 16°4’41”S, 145°27’13”E, 5–9.i.1983, ( T 54161View Materials in QM)GoogleMaps  ; Cape Tribulation transect Site 5, 16°4’41”S, 145°27’13”E, 23.ix –7.x.1982, ( T54185View Materials - T 54186View Materials in QM)GoogleMaps  ; Cape Tribulation transect Site 5A, 16°4’41”S, 145°27’0”E, 20–23.iv.1983, ( T54182View Materials T 54184View Materials in QM)GoogleMaps  ; Cape Tribulation transect Site 6, 16°4’41”S, 145°26’39”E, 23.ix-7.x.1982, ( T54167View Materials T 54177View Materials in QM)GoogleMaps  ; Cape Tribulation transect Site 6, 16°4’41”S, 145°26’39”E, 5–9.i.1983, ( T54178View Materials T 54180View Materials in QM)GoogleMaps  ; Cape Tribulation tran- sect Site 6, 16°4’41”S, 145°26’39”E, 20–23.iv.1983, ( T 54181View Materials in QM)GoogleMaps  ; Thompson Creek, ( Grove ), 16°6’’31”S, 145°26’25”E, 10.i–12.ii.1998, ( T73153View Materials T 73154View Materials in QM)  ; Thompson Creek, (Grove), 16°6’’31”S, 145°26’25”E, 15.iii.1998 – 7.v.1998, ( T 88806View Materials in QM)  ; Oliver Creek, 16°8’’22”S, 145°26’24”E, 4–7.vi.1974, ( T 49882View Materials in QM)  ; Cooper Creek, ( Grove ), 16°9’10”S, 145°24’19”E, 10.ii–16.iii.1998, ( T85646View Materials T 88801View Materials in QM)GoogleMaps  ; Pimm’s Block, ( Grove ), 16°11’20”S, 145°24’18”E, 8.i–9.ii.1998, ( T73155View Materials T 73160View Materials in QM)GoogleMaps  ; Pimm’s Block, ( Grove ), 16°11’20”S, 145°24’18”E, 9.ii–13.iii.1998, ( T85640View Materials T 85649View Materials in QM)GoogleMaps  ; Pimm’s Block, ( Grove ), 16°11’20”S, 145°24’18”E, 13.iii–8.v.1998, ( T88807View Materials T 88812View Materials in QM)GoogleMaps  ; Lync-Haven, Daintree area , 16°12’7”S, 145°24’32”E, 35 m, 2– 10.xii.2012, T.A. Weir & G.B. Monteith, (25-057656 1 specimen, 25-057658 4 specimens, 25-057662 3 specimens in ANIC)GoogleMaps  ; Lync-Haven, Daintree area , 16°12’7”S, 145°24’32”E, 35 m, 6–9.xii.2012, G.B. Monteith (25-057652 2 specimens, 25-057654 x 1 in ANIC)GoogleMaps  ; Lync-Haven, Daintree area , 16°12’7”S, 145°24’32”E, 35 m, 9.xii.2012, T.A. Weir (25-057665 3 specimens in ANIC)GoogleMaps  ; Mangrove Road, ( Grove ), 16°12’42”S, 145°25’18”E, 8.i-9.ii.1998, ( T73144View Materials T 73152View Materials in QM)GoogleMaps  ; Mangrove Road, ( Grove ), 16°12’42”S, 145°25’18”E, 9.ii–13.iii.1998, ( T 85642View Materials in QM)GoogleMaps  ; Mangrove Road, ( Grove ), 16°12’42”S, 145°25’18”E, 9.ix–25.x.1998, ( T88813View Materials T 88815View Materials in QM)GoogleMaps  ; Hutchinson Creek, ( Grove ), 16°13’4”S, 145°23’57”E, 13.i–11.ii.1998, ( T73161View Materials T 73167View Materials in QM)GoogleMaps  ; Hutchinson Creek, ( Grove ), 16°13’4”S, 145°23’57”E, 11.ii–14.iii.1998, ( T85643, T 85648 in QM)GoogleMaps  ; Hutchinson Creek, ( Grove ), 16°13’4”S, 145°23’57”E, 14.iii –8.v.1998, ( T88802View Materials T 88805View Materials in QM)GoogleMaps  ; Fairy Creek, ( Grove ), 16°14’16”S, 145°24’57”E, 11.ii– 13.iii.1998, ( T 85647View Materials in QM)GoogleMaps  ; Fairy Creek, ( Grove ), 16°14’16”S, 145°24’57”E, 13.iii–8.v.1998, ( T 88787View Materials in QM)GoogleMaps  ; Buchanan Creek , (Grove in QM)  , 16°14’39”S, 145°25’54”E, 5.i–11.ii.1998, ( T73141View Materials T 73142View Materials in QM)GoogleMaps  ; Buchanan Creek , (Grove in QM)  , 16°14’39”S, 145°25’54”E, 12.ix–23.x.1998, ( T88795, T 88797 in QM)GoogleMaps  ; Buchanan Creek , (Grove in QM)  , 16°14’39”S, 145°25’54”E, 12.iii–8.v.1998, ( T88796, T88798 T 88799View Materials in QM)GoogleMaps  ; Windsor Tableland via Mount Carbine , [16°15’0”S, 145°2’0”E], 25.xi.1976, R.I. Storey (25-059124 8 specimens in QDPC)GoogleMaps  ; Windsor Tableland ANZSES Site 9, 16°16’55”S, 145°3’56”E, 27.xii.1988 – 8.i.1989, ( T 54188View Materials in QM)GoogleMaps  ; Julatten (general), [16°37’0”S, 145°20’0”E], 21–30.xi.1987, A. Walford-Huggins (25-059114 5 specimens in ANIC)GoogleMaps  ; 6 km SW of Kuranda , [16°52’0”S, 145°37’0”E], 10.xii.1984 – 15.i.1985, R.I. Storey & H.K. Halfpapp, (25-059118 6 specimens in ANIC)GoogleMaps  ; 6 km SW of Kuranda , [16°52’0”S, 145°37’0”E], 15.i.1985 – 20.ii.1985, R.I. Storey & H.K. Halfpapp, (25- 032604 1 specimen in QDPC, 25-059117 5 specimens in QDPC)GoogleMaps  ; 6 km SW of Kuranda , [16°52’0”S, 145°37’0”E], 20.ii–20.iii.1985, R.I. Storey & H.K. Halfpapp, (25-059120 1 specimen in QDPC)GoogleMaps  ; 6 km SW of Kuranda , [16°52’0”S, 145°37’0”E], 6.xi–10.xii.1984, R.I. Storey & H.K. Halfpapp, (25-059119 4 specimens in QDPC)GoogleMaps  ; North Bell Peak summit, 17°5’19”S, 145°52’44”E, 15–16.ix.1981, ( T 54268View Materials in QM)GoogleMaps  ; Bellenden Ker Range, Cableway Base Station , [17°16’0”S, 145°55’0”E], 100 m, 17.x–9.xi.1981, Earthwatch, Queensland Museum, (25-059109 1 specimen in ANIC)GoogleMaps  ; Graham Range , summit, 17°16’24”S, 145°57’58”E, 1.xi–8.xii.1995, ( T 54240View Materials in QM)GoogleMaps  ; Graham Range , 350 m, 17°16’37”S, 145°57’48”E, 1.xi–8.xii.1995, ( T54237View Materials T 54239View Materials in QM)GoogleMaps  ; Wongabel State Forest , 6 km S of Atherton, [17°18’0”S, 145°31’0”E], 1.v–21.vi.1984, R.I. Storey & Brown (25-059126 1 specimen in ANIC)GoogleMaps  ; Topaz, Pei Road , (Frith house in QM)  , 17°24’23”S, 145°41’5”E, 6.xii.1993 – 25.ii.1994, ( T54242View Materials T 54244View Materials in QM)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 1.xii.1983 – 9.i.1984, R.I. Storey & Brown (25- 032603 1 specimen, 25-059111 4specimens in QDPC)GoogleMaps  ; Hugh Nelson Range , 21km S of Atherton, [17°25’0”S, 145°28’0”E], 1. v–21.vi.1984, R.I. Storey & Brown (25-059129 1 specimen in QDPC)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 10.ii–13.iii.1984, R.I. Storey & Brown (25-059127 2 specimens in QDPC)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 12.x–5.xi.1983, R.I. Storey & Brown (25-059131 1 specimen in QDPC)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 13.iii–1.v.1984, R.I. Storey & Brown (25-059128 3 specimens in QDPC)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 5.xi–1.xii.1983, R.I. Storey & Brown (25-059132 1 specimen in QCPC)GoogleMaps  ; Hugh Nelson Range , 21 km S of Atherton, [17°25’0”S, 145°28’0”E], 9.i–10.ii.1984, R.I. Storey & Brown (25-059133 1 specimen in QDPC)GoogleMaps  ; The Crater National Park , [17°25’0”S, 145°29’0”E], 16–18.iv.1973, R.I. Storey (25-058484 1 specimen in QM)GoogleMaps  ; Topaz, Hughes Road , (Breeden house in QM)  , 17°26’4”S, 145°42’2”E, 6.xii.1993 – 25.ii.1994, ( T 54241View Materials in QM)GoogleMaps  ; Millaa Millaa Falls , 17°29’53”S, 145°36’32”E, 6–9.ii.1999, ( T76200View Materials T 76201View Materials in QM)GoogleMaps  ; Millaa Millaa Falls (below), [17°29’0”S, 145°36’0”E], 760 m, 18–20.iv.1999, C. & I. Reid (25-059112 1 specimen in ANIC)GoogleMaps  ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 10.iv–4.v.1990, R.I. Storey & H.K. Halfpapp, (25- 032602 1 specimen in QDPC)GoogleMaps  ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 13.iii–10.iv.1990, R.I. Storey & H.K. Halfpapp, (25- 032601 3 specimens in QDPC)GoogleMaps  ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 4.i–7.ii.1990, R.I. Storey & H.K. Halfpapp, (25- 032599 8 specimens in QDPC)GoogleMaps  ; Millaa Millaa Falls , [17°30’0”S, 145°37’0”E], 760 m, 7.ii–13.iii.1990, R.I. Storey & H.K. Halfpapp, (25- 032600 4 specimens in QDPC)GoogleMaps  ; Kjellberg Road turnoff, 17°31’57”S, 145°36’27”E, 7.iii–15.v.1995, ( T54246View Materials T 54247View Materials in QM)GoogleMaps  ; Kjellberg Road turnoff, 17°31’57”S, 145°36’27”E, 10.i–31.iii.1995, ( T54248View Materials T 54250View Materials in QM)GoogleMaps  ; Mount Fisher, Kjellberg Road , 17°32’34”S, 145°33’31”E, 1.xii–25.ii.1994, ( T 54245View Materials in QM)GoogleMaps  ; Mount Fisher site BS2, [17°33’0”S, 145°32’0”E], 1150 m, 4.ix–3.x.1995, L. Umback (25-055628 1 specimen in ANIC)GoogleMaps  ; Maalan SF, Ravenshoe Road, 17°35’23”S, 145°35’40”E, 7.iii–15.v.1995, ( T54202View Materials T 54204View Materials in QM)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 10.i–31.iii.1995, ( T 54205View Materials in QM)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 10.i–31.iii.1995, ( T54206View Materials T 54208View Materials in QM)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 25.xi.1994 – 10.i.1995, ( T 54209View Materials in QM)GoogleMaps  ; Maalan State Forest, Ravenshoe Road , 17°35’23”S, 145°35’40”E, 6–9.ii.1999, ( T76197View Materials T 76199View Materials in QM)GoogleMaps  ; Mount Father Clancy , 900 m, 17°35’45”S, 145°38’4”E, 4.v.1983, T 54187View Materials in QMGoogleMaps  ); Rosina Creek  , 14 km SE Millaa Millaa , [17°36’0”S, 145°43’0”E], 720 m, 24.vi–2.viii.1982, S. & J. Peck (25-059107 1 specimen in ANIC)GoogleMaps  ; BS3View Materials Massey Creek , 17°37’0”S, 145°34’0”E], 1000 m, 1.xii.1994 – 3.i.1995, P. Zborowski 25-054102GoogleMaps  5 specimens in ANIC  ); BS3View Materials Massey Creek , 17°37’0”S, 145°34’0”E], 1000 m, 3.i–5.ii.1995, P. Zborowski (25-054106 1 specimen in ANIC)GoogleMaps  ; Plamerston National Park , [17°37’0”S, 145°47’0”E], 22–24.iv.1979, R.I. Storey (25-058483 1 specimen in QM)GoogleMaps  ; South Johnstone Forestry Camp , 5 km W, 17°38’12”S, 145°40’15”E, 1.xii.1993 – 25.ii.1994, ( T54211View Materials T 54214View Materials in QM)GoogleMaps  ; Charmillin Creek crossing, 17°42’3”S, 145°31’21”E, 8.xii.1989 – 5.i.1990, ( T54215View Materials T 54218View Materials in QM)GoogleMaps  ; Ravenshoe State Forest, Tully Falls Road, [17°43’0”S, 145°33’0”E], 10.x–15.xi.1987, A. Walford-Huggins (25-059113 1 in ANIC)GoogleMaps  ; Ravenshoe State Forest, Tully Falls Road, [17°43’0”S, 145°33’0”E], 15.xi–12.xii.1987, A. Walford-Huggins (25- 059115 1 in ANIC)GoogleMaps  ; Tully Falls State Forest 18 km SSW Ravenshoe, [17°46’0”S, 145°31’0”E], 730 m, 3.iii– 20.iv.1988, R.I. Storey & Dickinson, (25-059125 1 in QDPC)GoogleMaps  ; Tully Falls , 17°47’0”S, 145°33’27”E, 8.xii. 198 9– 5.i.1990, ( T 54220View Materials in QM)GoogleMaps  ; Tully Falls State Forest 11 km SSW Ravenshoe, [17°47’0”S, 145°34’0”E], 900 m, 1.x–5.xi.1987, R.I. Storey & Dickinson, (25-059116 2 specimens in QDPC)GoogleMaps  ; Tully Falls State Forest 11 km SSW Ravenshoe, [17°47’0”S, 145°34’0”E], 900 m, 5.i–7.ii.1988, R.I. Storey & Dickinson, (25-059122 2 specimens in QDPC)GoogleMaps  ; Tully Falls State Forest 11 km SSW Ravenshoe, [17°47’0”S, 145°34’0”E], 900 m, 5.xi–7.xii.1988, R.I. Storey & Dickinson, (25-059121 3 specimens in QDPC)GoogleMaps  ; Tully Falls State Forest 11 km SSW Ravenshoe, [17°47’0”S, 145°34’0”E], 900 m, 7.xii.1987 – 7.i.1988, R.I. Storey & Dickinson, (25-059123 2 specimens in QDPC)GoogleMaps  ; Boulder Creek track, 200–600 m, 17°51’2”S, 145°54’29”E, 24–27.x.1983, ( T54221View Materials T 54236View Materials in QM)GoogleMaps  .

Description. Holotype uniformly light brown. Others ranging from light brown to dark brown almost black. Antennal clubs grey.

Total length: 2.4–2.8 mm. Holotype measurements: total length 2.6 mm, elytral width 2.0 mm.

Male. Head. Surface very finely punctate, often appearing almost impunctate, basal carina absent. Clypeal edge normal, without projection lateral to teeth. Genal angle weakly defined. Dorsal part of the eyes narrow, separated by an interocular space approximately 20 times the eye width (40: 2); eye canthus nearly dividing eye. Head 1.3 x wider than long (48:36).

Prothorax: Pronotal surface with very fine punctation. Basal angles distinct. Punctures along the basal margin absent. Pronotum 1.8 x wider than long (41:23).

Elytra: Surface smooth with obsolete striae; intervals virtually impunctate. Striae 6 and 7 extending to elytral base. Elytra 1.1 x wider than long (51:45).

Legs: Protibiae usually with 3 teeth on outer edge, sometimes with only 2 teeth, front edge deeply angulate. Apical digit long and parallel sided with a rounded apex with a very small comb of stout setae, and bearing a short spur on the outer apical angle. Metatibiae with a notch on the inner edge.

Abdomen: Pygidium simple, entirely smooth, virtually impunctate and very slightly convex. Abdominal ventrites smooth, shiny and without reticulation. Ventrite 6 very finely punctate, almost impunctate.

Pterothorax: Medial lobe of metaventrite very finely punctate and very narrowly margined between the mesocoxae. Lateral lobe of metaventrite with inner border beside mesocoxae anteriorly curving to meet lateral border. Mesoventrite notched anteriorly. Mesoventrite smooth. Metanepisternum finely reticulate.

Aedeagus: Small tufts of setae projecting from apices of both right and left parameres, left paramere pointed.

Female. As per L. monteithi  species group. Metatibiae without notch on inner edge.

Etymology. Named in honour of Chris Reid, at the Australian Museum, for his invaluable contribution to the systematics of Australian beetles including the dung beetle genera Temnoplectron  and Coptodactyla  .

Distribution. Known from the rainforests of the Wet Tropics bioregion from Mount Boolbun south to Tully Falls National Park ( Fig. 12BView FIGURE 12).

Comments. Lepanus reidi  is known from almost 500 specimens, the majority of which have been collected in flight intercept traps or baited pitfall traps. This species appears to be a generalist and label data indicate this species has been collected in earthworm, cricket, or snail baited flight intercept traps, diurnally feeding on bird faeces, and from mushroom or dung baited pitfall traps (human or macropod faeces). Ebert et al. (2019) also collected L. reidi  new species from a pitfall trap baited with vertebrate carrion. A small number of specimens have been collected in malaise and window traps.

This species has been referred to under the informal nomenclature sensu Yeates et al. (2011) as Lepanus  NQ5 (Gunter et al. 2019). The specimen sequenced in this study was collected from the Mount Hypipamee National Park [formerly The Crater National Park] in the same trap as the holotype and is designated as a paratype here (ANIC 25- 060883 also as COL1152). It was recovered as sister to L. monteithi  in a larger clade containing all species within the L. monteithi  species group. Other specimens from Alexandra Range and Emmagen Creek have also been sequenced (25- 060882 and 25-057618). Lepanus reidi  and L. monteithi  are the only species in the group that have metatibiae with a notch on the inner edge and small tufts of setae on the apices of both parameres. Some specimens of L. reidi  can be interpreted as having only 2 teeth on the protibiae, but can be separated from L. monteithi  by the characters in the key.

ANIC

Australian National Insect Collection

QM

Queensland Museum

QDPC

Queensland Primary Industries Insect Collection

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Scarabaeidae

Genus

Lepanus