Diplomphalus Crosse & Fischer, 1872

Hausdorf, Bernhard, 2013, Revision of the endemic genera Diplomphalus and Pseudomphalus from New Caledonia (Gastropoda, Rhytididae), Zoosystema 35 (1), pp. 69-88: 70-84

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http://doi.org/ 10.5252/z2013n1a6

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http://treatment.plazi.org/id/03EBD306-B205-4138-C985-FA73E59E59F0

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scientific name

Diplomphalus Crosse & Fischer, 1872
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Genus Diplomphalus Crosse & Fischer, 1872  

Diplomphalus Crosse & Fischer, 1872: 288   . Type species (by monotypy): Helix cabriti Gassies, 1863   .

DIAGNOSIS. — Diplomphalus   is characterised by a small (4.4-9.0 mm) biconcave shell with a deeply sunken spire,

Revision of Diplomphalus   and Pseudomphalus   ( Gastropoda  , Rhytididae   ) from New Caledonia

tightly coiled whorls with an edge at the upper and the basal side and a very wide umbilicus.

Diplomphalus mariei (Crosse, 1867)   ( Figs 1 View FIG ; 2 View FIG ; 3A View FIG ; 4A, B View FIG ; 5 View FIG ; 6A, B View FIG )

Helix mariei Crosse, 1867a: 211   ; type locality: “Koe in vicinio urbis Noumea dictae, Novae-Caledonidae humi sub arbore reperta”, New Caledonia. — Crosse 1867b: 312, pl. 11 fig. 1. — Gassies 1871: 38, pl. 1 fig. 5.

Diplomphalus mariei   – Crosse & Fischer 1873: 23. — Tryon 1885: 113, pl. 25, figs 6, 7 (in part). — Kobelt 1904: 88, pl. 14 figs 3-5 (in part). — Franc 1957: 178, pl. 23, fig. 236 (in part).

Helix seberti Marie, 1881: 241   ; type locality: “ Kanala ”, New Caledonia. N. syn   .

Diplomphalus (Diplomphalus) mariei   – Crosse 1895: 175 (in part).

Diplomphalus (Diplomphalus) seberti   – Crosse 1895: 177, pl. 8, fig. 1.

Diplomphalus seberti   – Kobelt 1904: 89, pl. 14, figs 9-11. Franc 1957: 177, pl. 23, fig. 234.

Diplomphalus fischeri Franc, 1953: 81   , fig. 1; type locality: “ Nouvelle-Calédonie ”   ; 1957: 178, pl. 23, fig. 237. N. syn.

TYPE MATERIAL. — H. mariei: Koé   , 22°08’S, 166°28’E, 3 possible syntypes ( NHMUK 1883.11.10.842-844) GoogleMaps   ; H. seberti: Canala   , 3 syntypes ( MNHN; see also Crosse 1895: 177, pl. 8, fig. 1)   ; D. fischeri   : New Caledonia, many syntypes ( MNHN 22960-22962 View Materials ; see also Franc 1957: 178, pl. 23, fig. 237)   .

OTHER MATERIAL EXAMINED. — New Caledonia ( FMNH 36988 View Materials ; MNHN, among others voucher specimens to Franc 1957: 177, pl. 22, fig. 235 and p. 178, pl. 23, fig. 236; RBINS)   ; Forêt Plate, 540 m alt., 21°08’57”S, 165°06’23”E ( MNHN) GoogleMaps   ; Mt. Kopeto , 800 m alt., 21°10’39”S, 165°00’38”E ( MNHN) GoogleMaps   ; Adio, valley, 180 m alt., 21°14’44”S, 165°14’46”E ( MNHN) GoogleMaps   ; Adio , caves, 21°15’36”S, 165°14’18”E ( MNHN) GoogleMaps   ; Barrage de la Neaoua (Ouen Sieu), 500 m alt., 21°21’57”S, 165°32’35”E ( MNHN; ZMH 79086 View Materials ) GoogleMaps   ; Bogui, 150 m alt., 21°23’21”S, 165°26’30”E ( MNHN) GoogleMaps   ; Mines Emma-Pinpin, 50 m alt., 21°23’54”S, 165°16’51”E ( MNHN) GoogleMaps   ; Col des Roussettes , 200 m alt., 21°24’53”S, 165°26’44”E ( MNHN) GoogleMaps   ; Col des Roussettes , 270 m alt., 21°24’53”S, 165°26’44”E ( MNHN) GoogleMaps   ; Col des Roussettes , E side, Bourail-Houailou, 365 m alt. ( FMNH 158377 View Materials )   ; Bogui-Col des Roussettes , 270 m alt., 21°24’55”S, 165°26’49”E ( MNHN) GoogleMaps   ; Mé Aiu, 730 m alt., 21°29’15”S, 165°51’00”E ( MNHN) GoogleMaps   ; Koh, 150 m alt., 21°32’00”S, 165°48’50”E ( MNHN) GoogleMaps   ; Canala, 21°32’S, 165°57’E ( FMNH 73942 View Materials , 110159 View Materials ; MNHN; RBINS) GoogleMaps   ; Mé Ori, 530 m alt., 21°32’18”S, 165°40’22”E ( MNHN) GoogleMaps   ; Gelima, Caférie, 80 m alt., 21°32’24”S, 165°58’36”E ( MNHN) GoogleMaps   ; Mé Ori, 530 m alt., 21°32’37”S, 165°40’21”E ( MNHN) GoogleMaps   ; Table-Unio, 850-1000 m alt., 21°33’36”S, 165°45’55”E ( MNHN) GoogleMaps   ; Bourail , 21°34’S, 165°29’E ( FMNH 72860 View Materials ) GoogleMaps   ; 1.5 km inland from Pierced Rock, 8 km W of Bourail , 60 m alt. ( FMNH 158347 View Materials )   ; Dothio-Col de Pétchékara , 340 m alt., 21°34’32”S, 166°06’32”E ( MNHN) GoogleMaps   ; Col de Pétchékara , 450 m alt., 21°34’41”S, 166°05’48”E ( MNHN) GoogleMaps   ; Oua- Oué, 50 m alt., 21°37’00”S, 165°35’07”E ( MNHN) GoogleMaps   ; Col d’Amieu, summit, La Foa-Canala highway, 500 m alt., 21°37’S, 165°49’E ( FMNH 158417 View Materials , 158428 View Materials ) GoogleMaps   ; Nakada, 500 m alt., 21°38’57”S, 166°03’08”E ( MNHN) GoogleMaps   ; Farino , 21°40’S, 165°46’E ( RBINS) GoogleMaps   ; Fonwhary , 21°41’S, 165°47’E ( RBINS) GoogleMaps   ; Mine de Moindou , 21°42’S, 165°41’E ( FMNH 110158 View Materials ; RBINS) GoogleMaps   ; Mt. Ningua , 1000 m alt., 21°44’24”S, 166°09’03”E ( MNHN) GoogleMaps   ; Col de Nassirah , 250 m alt., 21°46’44”S, 166°04’14”E ( MNHN) GoogleMaps   ; Oua-Tom, 130 m alt., 21°47’15”S, 165°54’00”E ( MNHN) GoogleMaps   ; Oua Tom, 20 km N of Boulouparis , gully, 60 m alt., 21°48’S, 165°54’E ( FMNH 158400 View Materials , 158411 View Materials in part) GoogleMaps   ; Mont Mou, 400 m alt., 22°04’28”S, 166°19’46”E ( MNHN) GoogleMaps   ; Col de la Pirogue, Mont Mou, 450 m alt., 22°05’S, 166°20’E ” ( FMNH 144255 View Materials ) GoogleMaps   ; Monts des Koghis , 18 km N of Nouméa, 450 m alt. ( FMNH 158280 View Materials )   ; Nouméa , 22°16’S, 166°27’E ( MNHN) GoogleMaps   .

DISTRIBUTION ( FIG. 1 View FIG ). — Diplomphalus mariei   is restricted to the central part of New Caledonia. Its range overlaps those of D. solidula Tryon, 1885   and D. vaysseti ( Marie, 1871)   , demonstrating that it is specifically distinct from these species. At some localities it has been found even syntopically with D. vaysseti   . There is an old sample that is allegedly from Ouégoa (RBINS) in the northern part of the main island of New Caledonia where D. mariei   has not been found for more than a century, although several expeditions have collected there.

DIAGNOSIS. — Diplomphalus mariei   differs from D. solidula   and D. vaysseti   in a usually broader aperture (da/ha = 0.6-1.7 in D. mariei   , 0.5-0.7 in D. solidula   , 0.4-0.8 in D. vaysseti   ) and a callus forming an arcuate lamella projecting from the body whorl. The whorls of D. mariei   insert basally close to the edge of the previous whorl, whereas the whorls of D. solidula   and D. vaysseti   insert basally farther outside (and thus below) the edge of the previous whorl. Therefore the umbilical concavity of D. solidula   and D. vaysseti   is more stepped than that of D. mariei   .

DESCRIPTION

Shell ( Fig. 2 View FIG ; 3A View FIG ; Table 1) biconcave; spire deeply sunken, with 6.75-7 whorls with an edge at upper side, which can be compressed to form a keel, and an edge at basal side; protoconch initially smooth, then with microscopic spiral striae; teleoconch with moderately coarse, regular ribs (20-42 per quarter on upper side of body whorl) at upper and basal side that can split into 2 finer ribs at outer side of whorls between upper and lower edge and microscopic spiral threads; brownishcorneous with reddish-brown diagonal stripes; aperture pear-shaped, wider below than above; upper insertion of peristome distinctly descending towards aperture; peristome expanded, slightly reflexed, slightly thickened; insertions of peristome connected by a strong callus forming an arcuate lamella that projects from body whorl, especially in its middle; aperture without lamellae or folds; umbilicus wide.

Genitalia ( Fig. 4A, B View FIG ). Specimens from Col de Pétchékara and Col de Nassirah were examined. The genital pore is situated just posterior to the right ommatophore. The genitalia run to the right of the right ommatophoral retractor.Penis long and slightly broadened near the subterminal insertion of the vas deferens. There are straight longitudinal pilasters in proximal part of penis. Two strong longitudinal pilasters form a U-shaped structure. Opposite of the longitudinal pilasters, the penial wall is sculptured by wavy transversal pilasters. The narrow distal part of penis is sculptured with longitudinal pilasters. Penial retractor inserting terminally at penis and at diaphragm. Vas deferens divided in thin distal section running along penis towards atrium and thicker proximal section along vagina and free oviduct. Vagina shorter than free oviduct. Oviduct consisting of short narrower distal section and broader, longer glandular proximal section. Reservoir of bursa copulatrix only slightly broadened.

Mantle collar and pallial complex ( Fig. 5 View FIG ). Mantle collar broad (integument inserts deeply at the mantle edge), forming a narrow rim around the body without large lobes; only the region around the pneumostome is distinctly thickened; this thickening may form minute lobes at the edge. Heart about half as long as kidney, continuing anteriorly into indistinct pulmonary vein. Lung without additional coarse venation. Kidney long, opening anteriorly into reflexed primary ureter. Primary ureter running to the extreme posterior corner of the pulmonary cavity, where it transitions into the tubular secondary ureter, running along hindgut to the pnemostome, opening shortly before mantle collar, continuing anteriorly as a groove. There is no distinct mantle gland posterior to the mantle collar. Radula ( Fig. 6A, B View FIG ). Narrow; central teeth rudimentary, straight, stylet-like; 8 about equally long, slender, unicuspid lateral teeth on each side in V-shaped rows; basal half of lateral teeth attached with a small projection to the membrane, apical part curved towards the centre.

REMARKS

Helix seberti Marie, 1881   and Diplomphalus fischeri Franc, 1953   are synonymised here with Diplomphalus mariei   , because they fall into the range of variation of this species.

Diplomphalus solidula Tryon, 1885   ( Figs 4C View FIG ; 7 View FIG ; 8A View FIG )

Diplomphalus mariei   var. β – Crosse 1874: 387, pl. 12, fig. 1.

Diplomphalus mariei var. solidula Tryon, 1885: 114   , pl. 25, fig. 10; type locality: “Baie du Sud” (see Crosse 1874: 387), New Caledonia   .

Diplomphalus (Diplomphalus) mariei   – Crosse 1895: 175 (in part).

Diplomphalus (Diplomphalus) mariei   var. β Solidula – Crosse 1895: 176.

Diplomphalus mariei var. solidula   – Kobelt 1904: 89.

Diplomphalus mariei   – Franc 1957: 178 (in part, not Crosse 1867).

Diplomphalus vaysseti   – Franc 1957: 177, pl. 22, fig. 235 (in part, not Marie 1871).

TYPE MATERIAL. — D. mariei var. solidula: Baie du Sud   , syntype ( MNHN; voucher specimen of Crosse 1874: 387, pl. 12, fig. 1)   .

OTHER MATERIAL EXAMINED. — New Caledonia ( MNHN, among others the voucher specimen of Franc [1957: 177, pl. 22, fig. 235]; RBINS)   ; Baie du Sud ( MNHN; RBINS)   ; S of top of Mt. Ouin , 1150- 1200 m alt., 22°00’58”S, 166°28’21”E ( MNHN) GoogleMaps   ; Vallée Ouinné , 650 m alt., 22°02’23”S, 166°32’28”E GoogleMaps  

ps

( MNHN); Haute Rivière Bleue, 300 m alt., 22°04’07”S, 166°36’28’”E ( MNHN); Baie du Sud ( MNHN); Piste de la Pourina, 520 m alt., 22°05’00”S, 166°41’45”E ( MNHN); Mt. Kouvélè , E slope, 660 m alt., 22°05’26”S, 166°26’56”E ( MNHN); Rivière Bleue, 150 m alt., 22°05’47”S, 166°39’25”E ( MNHN); Rivière Bleue, 170 m alt., 22°05’59”S, 166°40’01”E ( MNHN); Rivière Bleue, 160 m alt., 22°06’13”S, 166°39’16”E ( MNHN); Parc à Cagous, Rivière Blanche, 160-170 m alt., 22°08’09”S, 166°38’20”E ( MNHN; ZMH 79089 View Materials ); near Mt. Ouenarou , 30 km NE of Nouméa (towards Yaté ), 250 m alt., 22°10’S, 166°43’E ( FMNH 158273 View Materials , 158361 View Materials ); Mont Guemba , 200 m alt., 22°10’22”S, 166°56’10”E ( MNHN); Mouirange , 180-250 m alt., 22°12’00”S, 166°39’00”E ( MNHN); Route les Dalmates, Monts Kouanenoa, E of St. Louis, 22°12’S, 166°42’E ” ( FMNH 144227 View Materials ) GoogleMaps   ; Mouirange , 200 m alt., 22°13’19”S, 166°40’14”E ( MNHN) GoogleMaps   ; S of Grand Lac , 280 m alt., 22°16’31”S, 166°54’00”E ( MNHN) GoogleMaps   ; Nouméa , Pointe de l’Artillerie, 22°17’S, 166°26’E ( RBINS) GoogleMaps   ; Forêt Nord , 200-250 m alt., 22°17’00”S, 166°53’00”E ( MNHN) GoogleMaps   ; Baie Ngo, 30 m alt., 22°18’02”S, 166°42’49”E ( MNHN; ZMH 79085 View Materials ) GoogleMaps   ; Baie du Prony ( RBINS)   ; Kwa Néie (Mts. Néngoné), 470 m alt., 22°19’09”S, 166°55’12”E ( MNHN) GoogleMaps   .

DISTRIBUTION ( FIG. 7 View FIG ). — Diplomphalus solidula   is restricted to the part of New Caledonia south of 22°S.

DIAGNOSIS. — Diplomphalus solidula   is characterised by a strong callus which extends a little forwards from the aperture and is slightly detached from the body whorl at its margin.

DESCRIPTION

Shell ( Fig. 8A View FIG ; Table 1) biconcave; spire deeply sunken, with 6.0-7.25 whorls with an edge at upper side and a weaker edge at the basal side; protoconch smooth; teleoconch with coarse, regular ribs (19-29 per quarter on the upper side of the body whorl) at upper and basal side that usually split into 2-3 finer ribs at outer side of whorls between upper and lower edge and microscopic spiral threads; brownish-corneous with reddish-brown transverse stripes; aperture unequally crescentic, wider below than above; upper insertion of peristome distinctly descending towards aperture; peristome not expanded, not or slightly reflexed, slightly thickened; insertions of peristome connected by a strong callus which extends a little forwards from aperture and is slightly detached from body whorl at its margin; aperture wuthout lamellae or folds; umbilicus wide. Genitalia ( Fig. 4C View FIG ). Specimens from Baie Ngo were examined. Penis long and broadened near subterminal insertion of vas deferens. There are straight longitudinal and wavy transversal pilasters inside the penis. Penial retractor inserts terminally at penis and at diaphragm.Vas deferens divided in thin distal section running along penis towards atrium and thicker proximal section along vagina and free oviduct. Vagina shorter than free oviduct. Oviduct consisting of short narrower distal section and broader, longer glandular proximal section.Reservoir of thin bursa copulatrix only slightly broadened.

Diplomphalus vaysseti ( Marie, 1871)   ( Figs 4D View FIG ; 7 View FIG ; 8B View FIG )

Helix volutella Gassies, 1858: 70   (not L. Pfeiffer, 1856); type locality: “bois de Balade”, New Caledonia. N. syn.

Helix Cabriti   – Gassies 1863: 21, pl. 1, fig. 4 (in part, not Gassies 1863).

Helix cabriti   – Crosse 1867b: 315 (in part, not Gassies 1863).

Helix vaysseti Marie, 1871: 325   ; type locality: “dans les montagnes, au cap Colnett”, New Caledonia   .

Diplomphalus cabriti   – Crosse & Fischer 1873: 22 (in part, not Gassies 1863). — Kobelt 1904: 87 (in part, not Gassies 1863). — Franc 1957: 176 (in part, not Gassies 1863).

Diplomphalus vaysseti   – Crosse & Fischer 1873: 23. — Crosse 1874: 388, pl. 12, fig. 2. — Tryon 1885: 113, pl. 25, figs 1-3. — Kobelt 1904: 89, pl. 14, figs 6-8. — Franc 1957: 177 (in part).

Helix vaysseti   – Gassies 1880: 33, pl. 1, fig. 21.

Diplomphalus (Diplomphalus) cabriti   – Crosse 1895: 173 (in part, not Gassies 1863).

Diplomphalus (Diplomphalus) vaysseti   – Crosse 1895: 176.

TYPE MATERIAL. — Unknown.

MATERIAL EXAMINED. — New Caledonia ( FMNH 36986 View Materials ; MNHN)   ; Col d’Amos, 200 m alt., 20°18’52”S,

Revision of Diplomphalus   and Pseudomphalus   ( Gastropoda  , Rhytididae   ) from New Caledonia

164°25’20”E ( MNHN)   ; Balade, 20°19’S, 164°30’E ( Gassies, 1858); 6.5 km E of Ouegoa , 120 m alt. ( FMNH 158250 View Materials , 159347 View Materials )   ; 8 km inland from Koumac , 90 m alt. ( FMNH 158300 View Materials )   ; Mandjelia, 400 m alt., 20°22’29”S, 164°30’06”E ( MNHN) GoogleMaps   ; Le Cresson, 20°29’00”S, 164°18’36”E ( MNHN) GoogleMaps   ; Cap Colnett, 20°30’S, 164°45’E ( Marie 1871); Kaala- Gomen, Konio, “la Cathédrale”, 350-450 m alt., 20°34’30”S, 164°24’12”E ( MNHN) GoogleMaps   ; Mont Panié near Tao ( FMNH 159344 View Materials )   ; Mont Panié, 280 m alt., 20°35’27”S, 164°48’43”E ( MNHN) GoogleMaps   ; Mont Panié, 580 m alt., 20°35’54”S, 164°48’22”E ( MNHN) GoogleMaps   ; Thiem, 10-50 m alt., 20°45’43”S, 165°06’23”E ( MNHN) GoogleMaps   ; Thiem , 90 m alt. ( FMNH 158326 View Materials )   ; valley SW of Oua Tilou , 200 m alt., 20°52’03”S, 164°49’49”E ( MNHN) GoogleMaps   ; Col Tango, way from Bobeitio, 300-350 m alt., 20°58’29”S, 165°00’27”E ( MNHN) GoogleMaps   ; Monéo, 21°09’36”S, 165°29’31”E ( MNHN) GoogleMaps   ; Mt. Aoupinié , 600-700 m alt., 21°10’47”S, 165°17’59”E ( MNHN) GoogleMaps   ; Adio, valley, 180 m alt., 21°14’44”S, 165°14’46”E ( MNHN) GoogleMaps   ; Adio, caves, 21°15’36”S, 165°14’18”E ( MNHN; ZMH 79087 View Materials ) GoogleMaps   ; 15 km NE of Poya, inland of Nékliaï , 150 m alt. ( FMNH 158358 View Materials )   ; Mé Maoya, 1250 m alt., 21°22’09”S, 165°20’21”E ( MNHN) GoogleMaps   ; Col des Roussettes , 200 m alt., 21°24’53”S, 165°26’44”E ( MNHN) GoogleMaps   ; Bogui-Col des Roussettes , 270 m alt., 21°24’55”S, 165°26’49”E ( MNHN) GoogleMaps   ; Table-Unio, 850-1000 m alt., 21°33’36”S, 165°45’55”E ( MNHN) GoogleMaps   .

DISTRIBUTION ( Fig. 7 View FIG ). — Diplomphalus vaysseti   is restricted to the northern part of New Caledonia. The gap between the distribution areas of D. vaysseti   and D. solidula   is not as large as shown in Figure 7 View FIG . There is a juvenile specimen from Oua Tom (FMNH 158411 in part) in the intermediate region which belongs to one of the two species.

Revision of Diplomphalus   and Pseudomphalus   ( Gastropoda  , Rhytididae   ) from New Caledonia

DIAGNOSIS. — Diplomphalus vaysseti   differs from D. solidula   and D. mariei   in a usually weak callus not detached from the body whorl and an on average denser coiling, i.e. a lower diameter at the same number of whorls. At five whorls the diameter of D. vaysseti   varies usually from 2.1-2.7 mm, in D. solidula   from 2.7-3.2 mm and in D. mariei   from 2.6-3.5 mm. D. vaysseti   differs from D. cabriti   in a smaller shell (diameter: 4.4-6.5 mm v 8.2-9.0 mm in D. cabriti   ).

DESCRIPTION

Shell ( Fig. 8B View FIG ; Table 1) biconcave; spire deeply sunken, with 6.5-8 whorls with an edge at upper side and an edge at basal side; protoconch initially smooth, then with microscopic spiral striae; teleoconch with coarse, regular ribs (20-36 per quarter on upper side of body whorl) at upper and basal side that can split into two finer ribs at outer side of whorls between upper and lower edge and microscopic spiral threads; brownish-corneous with a reticulate pattern of reddish-brown diagonal stripes; aperture unequally crescentic, wider below than above; upper insertion of peristome slightly descending towards aperture; upper part of peristome slightly protruding, lower part slightly reflexed, not expanded, slightly thickened; insertions of peristome connected by a weak callus which might extend a little forwards from aperture, but is not detached from body whorl; aperture without lamellae or folds; umbilicus wide.

Genitalia ( Fig. 4D View FIG ). Specimens from Col d’Amos and Bogui-Col des Roussettes were examined. Penis narrow and long and slightly broadened near subterminal insertion of vas deferens. There are straight longitudinal pilasters inside penis. Penial retractor inserts terminally at penis and at diaphragm. Vas deferens divided in thin distal section running along penis towards atrium and thicker proximal section along vagina and free oviduct. Vagina shorter than free oviduct.The oviduct consists of a short narrower distal section and a broader and longer glandular proximal section. Reservoir of bursa copulatrix only slightly broadened.

REMARKS

I could not examine type specimens of Helix volutella Gassies, 1858   (not L. Pfeiffer, 1856) and Helix cabriti Gassies, 1863   from Balade. However, only D. vaysseti   is known from this region of New Caledonia.The description and the measurements of H. volutella Gassies   fit D. vaysseti   . Thus, I consider this preoccupied name a synonym of D. vaysseti   . Helix cabriti Gassies   includes besides the specimens from Balade also specimens from the Île Art that belong to a different species that is treated in the following as D. cabriti   .

There are two specimens of D. vaysseti   from the northwest end of New Caledonia (Mandjelia and Le Cresson) which are 2.8 and 3.1 mm broad at 5 whorls, but have a weak callus and there are two specimens from Monéo which have a strong, detached callus, but which are only 2.3 mm broad at five whorls. These exceptions may be due to ancestral polymorphism, convergence or introgression between D. vaysseti   and D. solidula   .

Diplomphalus cabriti ( Gassies, 1863)   ( Figs 9 View FIG ; 10A View FIG )

Helix cabriti Gassies, 1863: 21   (in part); type locality (here restricted): “Île Art”, New Caledonia. — Crosse 1867b: 315, pl. 11, fig. 2 (in part). — Gassies 1871: 38 (?). — Fischer 1873: 9, pl. 3, fig. 8, 9.

Diplomphalus cabriti   – Crosse & Fischer 1872: 288. — Crosse & Fischer 1873: 22 (in part). — Tryon 1885: 113, pl. 25, figs 91, 92. — Kobelt 1904: 87, pl. 13, figs 21, 22 (in part). — Franc 1957: 176, pl. 23, fig. 233 (in part). — Schileyko 2000: 752, fig. 980.

Diplomphalus (Diplomphalus) cabriti   – Crosse 1895: 173 (in part). — Wenz & Zilch 1960: 551, fig. 1926.

TYPE MATERIAL. — Île Art, lecto- and paralectotype ( NHMUK 1883.11.10.289, 290).

OTHER MATERIAL EXAMINED. — New Caledonia ( FMNH 36985; MNHN, among others voucher specimen of

Crosse [1867b: 315, pl. 11, fig. 2]); Île Art ( MNHN; RBINS).

DISTRIBUTION ( FIG. 9 View FIG ). — Diplomphalus cabriti   is known only from the Île Art without more exact locality. A record from Pouébo in the northernmost part of the main island of New Caledonia ( Gassies 1871; Crosse & Fischer 1873) is probably erroneous or might refer to Diplomphalus vaysseti   (see also D. montrouzieri ( Souverbie, 1858))   .

DIAGNOSIS. — Diplomphalus cabriti   differs from all other Diplomphalus   species in a larger shell (diameter: 8.2-9.0 mm v 4.4-7.9 mm in all other species).

DESCRIPTION

Shell ( Fig. 10A View FIG ; Table 1) biconcave; spire deeply sunken, with 8.75-9.75 whorls with a compressed keel at upper side and an edge at basal side; protoconch smooth; teleoconch with coarse, regular ribs (26-39 per quarter on upper side of body whorl) at upper and basal side that usually split into 2-3 finer ribs at outer side of whorls between upper and lower edge and microscopic spiral threads; corneous with a reticulate pattern of reddish-brown diagonal stripes; aperture narrow crescentic, slightly wider below than above; upper insertion of peristome distinctly descending towards aperture; peristome slightly expanded, not reflexed, slightly thickened; insertions of peristome connected by a weak callus which is thickened at its margin, but not detached from body whorl; aperture without lamellae or folds; umbilicus wide.

Genitalia unknown.

Radula. Fischer (1873) counted 16 about equally long, slender, unicuspid lateral teeth on each side. He did not find central teeth, but if they are as small as in D. mariei   , he might have missed them.

REMARKS

Gassies (1863) included specimens from Balade probably belonging to Diplomphalus vaysseti   (see above) as well as specimens from the Île Art in his Helix cabriti   . All later descriptions and figures of D. cabriti   refer to the species from the Île Art. I could not examine type specimens of H. cabriti   from Balade. However, there are two specimens from the Île Art labelled as types of H. cabriti Gassies   in the Natural History Museum in London (NHMUK 1883.11.10.289-290). I designate one of these specimens (the one glued at the basal side to the card board; measurements shell diameter: 9.1 mm, shell height: 4.2 mm) as lectotype of H. cabriti Gassies, 1863   to clarify the taxonomic status of the name and to preserve stability of nomenclature.

Diplomphalus cabriti   is probably most closely related with D. montrouzieri   , which is also larger than the Diplomphalus   species from the main island of New Caledonia and which also has a callus which is thickened at its margin, but is not detached from the body whorl.

Diplomphalus montrouzieri ( Souverbie, 1858)   ( Figs 9 View FIG ; 10B View FIG )

Helix montrouzieri Souverbie, 1858: 63   ; type locality: “Artensis insula”, New Caledonia. — Fischer 1858: 296, pl. 8, fig. 7. — Gassies 1863: 20, pl. 1, fig. 3. — Gassies 1871: 38.

Diplomphalus montrouzieri   – Crosse & Fischer 1873: 23. — Tryon 1885: 113, pl. 25, fig. 90. — Kobelt 1904: 87, pl. 14, figs 1, 2. — Franc 1957: 177.

Diplomphalus (Diplomphalus) montrouzieri   – Crosse 1895: 175.

TYPE MATERIAL. — Île Art, 3 syntypes ( MNHN).

OTHER MATERIAL EXAMINED. — New Caledonia ( FMNH 36983 View Materials , 36984 View Materials ; MNHN)   ; Île Art ( MNHN, among others 3 syntypes of H. montrouzieri Souverbie   ; RBINS; ZMH 79088 View Materials )   ; “Pouébo” ( MNHN); “Canala” ( RBINS).

DISTRIBUTION ( FIG. 9 View FIG ). — Diplomphalus montrouzieri   is probably restricted to the Île Art. There is a sample from Pouébo in the northern part of the main island of New Caledonia (MNHN) from where D. montrouzieri   and also D. cabriti   have been recorded by Gassies (1871). However, neither of these species has been found in that region of the main island of New Caledonia for more than a century, although several expeditions have collected there. There is also a sample that is allegedly from Canala (RBINS), but this is almost certainly an error.

DIAGNOSIS. — Diplomphalus montrouzieri   is larger than the Diplomphalus   species from the main island of New Caledonia, but smaller than D. cabriti   (diameter of D. montrouzieri   : 6.7-7.9 mm; of D. cabriti   : 8.2- 9.0 mm), from which it differs also in a whitish shell colour without brownish pattern and a weaker sculpture of the teleoconch.

DESCRIPTION

Shell ( Fig. 10B View FIG ; Table 1) biconcave; spire deeply sunken, with 8.0-8.50 whorls with a compressed edge at upper side and an edge at basal side; protoconch smooth; teleoconch with flat, regular ribs (21-31 per quarter on upper side of body whorl) at upper and basal side that usually split into 2-3 finer striae at outer side of whorls between upper and lower edge and microscopic spiral threads; whitish; aperture narrow crescentic; upper insertion of peristome distinctly descending towards aperture; peristome not expanded, not reflexed,not thickened; insertions of peristome connected by a weak callus thickened at its margin, but not detached from body whorl; aperture without lamellae or folds; umbilicus wide. Genitalia unknown.

NHMUK

Natural History Museum, London

MNHN

Museum National d'Histoire Naturelle

RBINS

Royal Belgian Institute of Natural Sciences

ZMH

Zoologisches Museum Hamburg

FMNH

Field Museum of Natural History

Kingdom

Animalia

Phylum

Mollusca

Class

Gastropoda

Order

Stylommatophora

Family

Rhytididae

Loc

Diplomphalus Crosse & Fischer, 1872

Hausdorf, Bernhard 2013
2013
Loc

Diplomphalus mariei

FRANC A. 1957: 178
1957
Loc

Diplomphalus vaysseti

FRANC A. 1957: 177
1957
Loc

Diplomphalus fischeri

FRANC A. 1953: 81
1953
Loc

Diplomphalus seberti

FRANC A. 1957: 177
KOBELT W. 1904: 89
1904
Loc

Diplomphalus mariei var. solidula

KOBELT W. 1904: 89
1904
Loc

Diplomphalus (Diplomphalus) mariei

CROSSE H. 1895: 175
1895
Loc

Diplomphalus (Diplomphalus) seberti

CROSSE H. 1895: 177
1895
Loc

Diplomphalus (Diplomphalus) mariei

CROSSE H. 1895: 175
1895
Loc

Diplomphalus (Diplomphalus) mariei

CROSSE H. 1895: 176
1895
Loc

Diplomphalus (Diplomphalus) cabriti

CROSSE H. 1895: 173
1895
Loc

Diplomphalus (Diplomphalus) vaysseti

CROSSE H. 1895: 176
1895
Loc

Diplomphalus (Diplomphalus) cabriti

WENZ W. & ZILCH A. 1960: 551
CROSSE H. 1895: 173
1895
Loc

Diplomphalus (Diplomphalus) montrouzieri

CROSSE H. 1895: 175
1895
Loc

Diplomphalus mariei var. solidula

TRYON G. W. 1885: 114
CROSSE H. 1874: 387
1885
Loc

Helix seberti

MARIE E. 1881: 241
1881
Loc

Helix vaysseti

GASSIES J. - B. 1880: 33
1880
Loc

Diplomphalus mariei

CROSSE H. 1874: 387
1874
Loc

Diplomphalus mariei

FRANC A. 1957: 178
KOBELT W. 1904: 88
TRYON G. W. 1885: 113
CROSSE H. & FISCHER P. 1873: 23
1873
Loc

Diplomphalus cabriti

FRANC A. 1957: 176
KOBELT W. 1904: 87
CROSSE H. & FISCHER P. 1873: 22
1873
Loc

Diplomphalus vaysseti

FRANC A. 1957: 177
KOBELT W. 1904: 89
TRYON G. W. 1885: 113
CROSSE H. 1874: 388
CROSSE H. & FISCHER P. 1873: 23
1873
Loc

Diplomphalus montrouzieri

FRANC A. 1957: 177
KOBELT W. 1904: 87
TRYON G. W. 1885: 113
CROSSE H. & FISCHER P. 1873: 23
1873
Loc

Diplomphalus

CROSSE H. & FISCHER P. 1872: 288
1872
Loc

Diplomphalus cabriti

SCHILEYKO A. A. 2000: 752
FRANC A. 1957: 176
KOBELT W. 1904: 87
TRYON G. W. 1885: 113
CROSSE H. & FISCHER P. 1873: 22
CROSSE H. & FISCHER P. 1872: 288
1872
Loc

Helix vaysseti

MARIE E. 1871: 325
1871
Loc

Helix mariei

GASSIES J. - B. 1871: 38
CROSSE H. 1867: 211
CROSSE H. 1867: 312
1867
Loc

Helix cabriti

CROSSE H. 1867: 315
1867
Loc

Helix

GASSIES J. - B. 1863: 21
1863
Loc

Helix cabriti

FISCHER P. 1873: 9
GASSIES J. - B. 1871: 38
CROSSE H. 1867: 315
GASSIES J. - B. 1863: 21
1863
Loc

Helix volutella

GASSIES J. - B. 1858: 70
1858
Loc

Helix montrouzieri

GASSIES J. - B. 1871: 38
GASSIES J. - B. 1863: 20
SOUVERBIE M. 1858: 63
FISCHER P. 1858: 296
1858