Tylopus punctus, Likhitrakarn & Golovatch & Panha, 2016

Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak, 2016, The millipede genus Tylopus Jeekel, 1968 (Diplopoda, Polydesmida, Paradoxosomatidae), with a key and descriptions of eight new species from Indochina, European Journal of Taxonomy 195, pp. 1-47 : 29-32

publication ID	https://doi.org/ 10.5852/ejt.2016.195
DOI	https://doi.org/10.5281/zenodo.3852406
persistent identifier	https://treatment.plazi.org/id/03EB5A00-FFE3-7D77-ACE7-D63DFE1E990C
treatment provided by	Valdenar
scientific name	Tylopus punctus
status	sp. nov.

Treatment

Tylopus punctus View in CoL sp. nov.

urn:lsid:zoobank.org:act:35585ACA-8490-49EE-8DA5-73EB3EB04A5C

Figs 21–22 View Fig View Fig , 26 View Fig

Diagnosis

Very similar to T. silvestris ( Pocock, 1895) , but differs in the smaller size, the width of midbody being 2.1–2.5 mm (versus 3.0 mm); the caudal corner of midbody paraterga very narrowly rounded and not drawn behind the rear tergal margin (versus nearly pointed and produced behind the rear tergal margin), and gonopod process h small and pointed (versus absent).

Etymology

To emphasize gonopod process h being small and pointed; adjective.

Material examined

Holotype

MYANMAR: Ƌ, Mintaingbin Forest Camp , ca 35 km north of Aungban, Chan State, 20°55'20" N, 96°33'60" E, ca 1300 m a.s.l., 11–23 Jun. 2004, leg. H. Schillhammer & H. V. Shaverdo (NHMW- 8248).

GoogleMaps

Paratype

MYANMAR: 1 ♀, same data as for holotype (NHMW-8249).

Description

MEASUREMENTS AND COLOUR. Length 27.5 (Ƌ) or 29 mm (♀), width of midbody pro- and metazonae 1.56 and 2.1 mm (Ƌ) or 1.98 and 2.5 mm (♀), respectively. Coloration of alcohol material upon long-term preservation rather uniformly light brown to brown, with light yellow paraterga, antennae, venter and legs ( Fig. 21 View Fig A–I).

HEAD. Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae rather short ( Fig. 21B View Fig ), reaching only in front of body segment 3 (Ƌ, ♀) when stretched dorsally. In width, segment 3 <4 <head <collum <segment 2 <5–15; thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 4+4 anterior, 2+2 intermediate and 3+3 posterior; an evident lateral incision at about anterior 1/3; caudal corner of paraterga very broadly rounded, declined ventrad, not drawn behind rear tergal margin.

BODY. Tegument smooth and shining, prozonae very finely shagreened, metaterga rather smooth and leathery, with posterior halves faintly rugulose, surface below paraterga finely microgranulate ( Fig. 21 View Fig A–F). Postcollum metaterga with two transverse rows of setae: 2+ 2 in anterior (pre-sulcus) row, mostly abraded, and 2+ 2 in posterior (post-sulcus) row, mostly traceable as insertion points, except for segments 18 and 19, each latter with 4+4 setae in posterior row. Tergal setae simple, slender, about 1/3 of metatergal length ( Fig. 21A, F View Fig ). Axial line visible both on pro- and metazonae. Paraterga welldeveloped ( Fig. 21 View Fig A–G), especially so in Ƌ, set high (at upper ¼ of body height), anterior edge rounded and narrowly bordered, fused to callus; caudal corner very narrowly rounded, segments 2 and 3 slightly extending behind rear tergal margin, thereafter not surpassing rear tergal margin until segment 15 ( Fig. 21 View Fig A–D). Paratega 2 broad, anterior edge angular, lateral edge with two small, but evident incisions in anterior half. Following paraterga with an evident incision in anterior 1/3 of lateral edge ( Fig. 21A, C, F View Fig ). Calluses on paraterga delimited by a sulcus only dorsally, rather narrow, a little wider on porebearing segments. Ozopores distinct, lying at about ¼ of metatergal length in front of posterior edge of metaterga. Transverse sulcus usually distinct ( Fig. 21A, D, F View Fig ), slightly incomplete on segment 19, complete on segments 5–18, narrow, line-shaped, rather deep, reaching bases of paraterga, beaded at bottom. Stricture between pro- and metazonae evident, rather wide and deep, beaded at bottom down to base of paraterga ( Fig. 21A View Fig , C–F). Pleurosternal carinae complete crests with a sharp caudal tooth on segments 2 and 3, thereafter increasingly reduced and remaining a sharp caudal tooth until segment 9 (Ƌ) or 6 (♀), retained as a small caudal tooth and increasingly reduced until segment 17, absent from segment 18 (Ƌ) or 7 (♀). Epiproct ( Fig. 21 View Fig E–G) conical, flattened dorsoventrally, with two evident, rather large, rounded, apical papillae; tip subtruncate; lateral pre-apical papillae clear, lying close to tip. Hypoproct roundly subtriangular, setigerous knobs at caudal edge small and well-separated ( Fig. 21G View Fig ).

STERNA. Sparsely setose, without modifications, but with a pair of small, blunt, fully separated cones between Ƌ coxae 4 ( Fig. 21H, I View Fig ). Legs rather short and slender, midbody ones ca 1.1–1.2 (Ƌ) or 0.8–0.9 (♀) as long as body height, Ƌ prefemora not swollen, Ƌ tarsal brushes present only on legs 1 and 2.

GONOPOD. Rather simple ( Fig. 22 View Fig ); coxa slightly curved caudad, sparsely setose distoventrally. Prefemur sparsely setose, about 1/3 as long as femorite + postfemoral part. Femorite curved and stout, with an evident mesal groove and a strong distolateral sulcus demarcating a postfemoral part; process h small, slender and pointed; solenophore (sph) clearly coiled, long and flattened.

Remark

Pocock (1895) described only T. doriae ( Pocock, 1895) and T. silvestris from northern Myanmar, the above being a third species of this genus also occurring in northern Myanmar.

New faunistic record