Coremiocnemis hoggi, West & Nunn, 2010

West, Rick C. & Nunn, Steven C., 2010, 2443, Zootaxa 2443, pp. 1-64 : 28-46

publication ID

1175­5334

persistent identifier

https://treatment.plazi.org/id/03EAD663-FFF5-FF84-FACE-F8C2FEE2E879

treatment provided by

Felipe

scientific name

Coremiocnemis hoggi
status

sp. nov.

Coremiocnemis hoggi View in CoL sp. nov.

( Figs 78–118, 217)

Types: female holotype, S88042 View Materials , Fraser’s Hill, Selangor, West Malaysia, col. R. West, 30 Mar. 2001, QM – examined; male paratype, S88022 View Materials , Fraser’s Hill, Selangor, West Malaysia, col. S. Yong, 24 Sept. 2008, QM – examined; female paratype, S74074 View Materials , Fraser’s Hill, Selangor, West Malaysia, col. R. West and J. Huff, 10 April 2000, QM –examined; 2 females S88076 View Materials , S88078 View Materials , Fraser’s Hill, Selangor, West Malaysia, col. S. Hogg, 22 Feb. 2009, QM –examined; female paratype, ZRC ARA.741, Fraser’s Hill, Selangor, West Malaysia, col. E. Loh and K. Lim, 2 June 1990, RMBR –images (D. Court, pers. comm.)–examined; female paratype, ZRC ARA.67, Fraser’s Hill, Selangor, West Malaysia, col. H. M. Pendlebury, 1 Feb. 1929, RMBR –images (D. Court, pers. comm.)–examined; 2 female paratypes, ZRC ARA.756, ZRC ARA.757, Fraser’s Hill , Selangor, West Malaysia, ( Gopal series, 1986–89), RMBR –images (D. Court, pers. comm.)–examined .

Other material: 2 juveniles, S88077 View Materials , S88079 View Materials , Fraser’s Hill, Selangor, West Malaysia, col. S. Hogg, 22 Feb. 2009, QM –examined .

Etymology: A patronym in honor of Mr Stephen Hogg who gathered specimens and natural history data on Coremiocnemis species from Fraser’s Hill and Maxwell Hill, West Malaysia, for this paper.

Diagnosis: Differs from all other Coremiocnemis species in having extremely long setae on leg IV (older adult females), almost entire scopula on retrolatero-ventral metatarsi IV and leg IV thickest cf. I–III. Differs from C. cunicularia in lacking distinct white/cream bands along the distal segments of all legs (distinct and obvious in C. cunicularia ), spermathecal morphology, the colour of femoral leg segments and the lyra bacillae (shafts 1.5 times thicker than those of C. cunicularia ). Differs from C. gnathospina sp. nov. in the reduced number of intercheliceral pegs (under>45). Differs from C. valida in lacking ventral endocuticular black studs bordering the sternum and coxae III and IV, and scopula in only the distal 1/3 metatarsi IV ventrally on retrolateral side (entire retrolaterally in C. valida ). Differs from C. kotacana sp. nov., C. brachyramosa sp. nov., C. jeremyhuffi sp. nov. and C. obscura sp. nov. in the light setal brush along the median retrolateral surface of coxae IV (surface pallid in aforementioned species).

Description: Female holotype QM S88042 View Materials ( Fig. 78) with body length: 53.73.

Color (in life, Fig. 78): Carapace, chelicerae, coxae and trochantera dark brown (abdomen only slightly darker), anterior distal leg segments (pat., tib., met., tar.) light sandy. Femora dark with strong violet hue in natural light. Ventrally dark brown/black. All leg segments lack cream banding distally. Long setae on posterior legs same color as anterior legs, light sandy.

Carapace ( Fig. 80): length 20.48, width 17.54 (width across anterior edge 11.48). Fovea 2.81 in width, procurved, deep, not quite as wide as OT ( Fig. 81).

Eyes ( Fig. 81): ocular tubercle; length 2.15, width 3.31. Anterior almost straight, slightly procurved. Posterior row slightly recurved. Eyes: ALE, AME, PLE, PME.

Chelicerae ( Figs 93–95): length 10.95, width at base of each lobe 4.99. Ectal lyrate region a series of strikers (>50) in>7 horizontal rows ( Fig. 94). Strongest, longest strikers on lowest rows. Each striker needleform, with some filiform ends. Teeth 15 (right),>100 basomesals. Intercheliceral pegs (>4 larger,>18 smaller) in tight cluster on basodorsal surface ( Fig. 95). On retrolateral face basomedially, small cluster of short strong spines (>3), point distad.

Maxillae ( Figs 82, 96): prolaterally plano-convex, anterior lobe well pronounced, many cuspules (> 200) on inner basoventral surface. Lyra ( Fig. 96): manybacilliform rods (>200) in dense, ovoid patch on prolateral maxillary surface, lowest row with>30 bacillae, longest rods medially in lowest row. Rods paddle-shaped with medium to long shafts, largest with distal blades. At widest point, lyrate patch is>10 rows deep with smallest rods dorsally. Posterior end of patch truncate, anterior end slightly pointed distad/rounded. Immediately above maxillary suture>55 small spines on anterior margin of maxillae, rows unordered.

Labium ( Fig. 83): length 2.89, width 3.62. Many small cuspules (>500) along anterior 1/3 surface.

Sternum ( Fig. 84): length 9.98, width 9.34. Slightly wider posteriorly than anteriorly, with thin but even cover of short setae, interspersed with longer darker setae. No spines along sternal border, some strong spiniform setae on lateral sternal margins. Posteriorly between left–right cox. IV, sternal border slightly acuminate, lateral sternal points slightly acuminate. Sigilla ( Fig. 84): 3 pairs (not including labio-sternal sigilla), posterior medium, right with length/width:1.43/0.69, left 1.56/0.78 ovoid, 2.65 apart and 1.75–1.96 from sternal margin. Median pair 1/2 size of posterior, similar in form, 1.2–1.4 of their length from sternal margin. Anterior pair somewhat obscured, in sternal margin. Labio-sternal sigilla small, thin, but wide.

Leg setation ( Figs 86–90): Leg IV with long curling setae on all segments, most noticeable on tib. and met. IV, having “bottlebrush” appearance. Entirely on tib. and met. IV, longer setae show distinct recurvature (most obvious in young adults–juveniles). In older specimens, long setae become elongate (losing recurvature of setae on tib. IV), having extreme bottlebrush appearance. Long recurved setae distinct retrolaterally on tar. IV.

Legs: formula; length IV, I, II, III; width IV, III, I, II. Leg RF ~ 80.39. Leg lengths (fem., pat., tib., met., tar., total): holotype QM S88042 View Materials : palp: 11.11, 6.78, 7.9, 0.00, 6.91, 32.7. I: 16.51, 10.6, 12.65, 10.99, 5.29, 56.04. II: 14.17, 8.56, 10.18, 9.84, 5.84, 48.59. III: 13.61, 8.53, 8.93, 10.37, 5.63, 47.07. IV: 19.17, 10.14, 15.08, 18.68, 6.64, 69.71. Leg mid-widths (fem., pat., tib., met., tar., total.): palp: 3.13, 2.91, 2.76, 0.00, 2.68, 11.48. I: 3.27, 3.58, 2.99, 2.71, 2.66, 15.21. II: 3.47, 3.63, 2.81, 2.21, 2.34, 14.46. III: 4.23, 4.12, 3.27, 2.46, 2.51, 16.59. IV: 4.59, 4.06, 3.51, 3.14, 2.46, 17.76.

Scopula: Met. and tar. I–III undivided; met. and tar. IV divided ( Fig. 87). Met. I, II: entire, III: 5/6, IV: 5/6 retrolateral side of division; 4/5 prolaterally ( Figs 86–88).

Coxae: some small black thorns prolatero-dorsally, no thorns retrolaterally on I–IV. Coxae easily seen dorsally. I longest, ca. 1.3 times length of II. IV widest, longer than III, basally rectangular with well defined corners. Coxae largely lacking ventral thorns prolaterally on I–IV (strong dark setae only). I–III ventrally with many long thick blunt setae proximally, pallid. No short black setae. IV with mixture of long thick blunt setae proximally, longer wavy setae along retrolateral ledge, pallid intermixed with shorter thin pallid setae. Ventral I–IV gently sloping anteriorly. Retrolateral setation: I–III with median narrow light brush, IV as in I–III, with light narrow brush. I–IV retrolaterally with defined ventral ledge. Ventral measurements for coxae (QM S88042 View Materials ): palp–length 10.59, width 5.39; I–9.82, 4.29; II–7.6, 4.23; III–7.86, 4.51; IV–9.48, 4.91.

Trochantera: palp–length 2.74, width 2.86; I–4.0, 4.12; II–3.1, 3.89; III–3.11, 4.06; IV–3.85, 4.40.

Trichobothria: on all tarsi basal filiform field slightly wider than clavate field, merges evenly. Clavates on tarsi I in distal 2/3, long filiforms only in basal 1/2, shorter filiforms intermixed with clavates distally. Clavate extent on II–IV cf. I, in distal 2/3. Shorter filiforms for length. Short epitrichobothrial field on I shorter than clavate, increasing in length proximally.

Tarsal organ evident on legs I–IV, low domed, but small ( Figs 91, 92).

Spines: met. I with 1 DV, met. II with 1 DV, 1 DPV, 1 DRV, met. III with 1 DV, 1 DPV, 1 DRV, 1 DD, 1 DPL and met. IV with 1 DV, 1 DPV, 1 DRV, 1 DD, 1 DPL.

Claws: paired claws on all legs and palpal claw unarmed. Reduced third claw on leg IV.

Abdomen ( Fig. 85): length 23.23, width 14.04. Oval, elongated, covered with mat of very fine short dark brown setae with sparse cover of longer red/brown setae. Longer hairs concentrated ventrally toward spinnerets, point distad. Slightly darker near booklung–epigynum areas.

Genitalia ( Fig. 84): epigastric fold 4.68 wide. Spermathecae paired, not fused, bilobular apically with each lobe with strong apical rounded appendages, heavily sclerotized compared to spermathecal shafts, sclerotization reducing gradually. Shafts basally incrassate, tapering distally to swollen tips. Lobes apically swollen, lateral lobes as long as medials. Epigastric fold extends ca. 2/3 length of lobes.

Spinnerets: PMS: length 2.42, width (medially) 0.89. PLS: section lengths; basal 4.01, medial 2.43, apical 3.78. PLS section widths (medially); basal 1.85, medial 1.51, apical 1.12.

Male paratype QM S88022 View Materials (unless otherwise stated and as in female, except) with body length: 27.60.

Color (in life, Fig. 79): Carapace, coxae, trochantera with strong pinkish hue in natural light. Femora jet black with long wavy pale setae giving speckled appearance. Abdomen, distal leg segments (pat., tib., met., tar.) pale sandy brown (white in postmoult individuals).

Carapace ( Fig. 97): length 11.65, width 9.91 (width across anterior edge 5.49). Fovea 1.34, deep.

Eyes ( Fig. 98): ocular tubercle; length 1.56, width 2.21. Anterior row transverse. Posterior row mildly recurved, almost transverse. Eyes: AME, ALE, PLE, PME.

Chelicerae ( Figs 103–107): length 7.04, width at base of each lobe 2.54. Teeth 11,>65 smaller basomesals. Intercheliceral pegs (>3 larger,>15 smaller) in tight cluster on basodorsal surface ( Fig. 106).

Maxillae ( Figs 99, 101): anterior lobe well pronounced,>160 cuspules on inner basoventral surface. Maxillary lyra ( Fig. 102): dorsal to maxillary suture 18 to 25 elongated spines medially to anterior margin of maxillae. Bacillate rods slightly shorter than in female, 10–16 with long shafts. Longest bacillae with distal blades, up to 3 times length cf. females.

Labium ( Fig. 100): length 1.44, width 2.01. At least 550 small cuspules along anterior 1/3.

Sternum ( Fig. 99): length 4.84, width 5.09. Clearly wider posteriorly than anteriorly (but not as elongate as females), with thin but even cover of short setae, interspersed with many longer darker setae. No spines on sternal border. Lateral sternal points acuminate, posterior point not.

Legs: formula; length IV, I, II, III; width IV, I, II, III. Leg RF ~ 95.51. Leg lengths (fem., pat., tib., met., tar., total): palp: 7.30, 5.07, 7.22, 0.00, 2.22, 21.81. I: 10.18, 5.92, 10.52, 8.23, 4.93, 39.78. II: 9.46, 5.24, 7.98, 7.95, 4.56, 35.19. III: 7.36, 4.42, 6.27, 7.74, 4.20, 29.99. IV: 10.48, 4.91, 9.44, 12.53, 4.29, 41.65. Leg mid– widths (fem., pat., tib., met., tar., total.): palp: 1.69, 1.99, 1.77, 0.00, 2.02, 7.47. I: 2.21, 2.33, 1.93, 1.44, 1.45, 9.36. II: 2.31, 2.11, 1.81, 1.46, 1.46, 9.15. III: 2.89, 2.30, 1.59, 1.19, 1.06, 9.03. IV: 2.30, 2.27, 2.15, 1.29, 1.22, 9.23. Coxae ventral measurements: palp–length 5.01, width 2.17; I–5.06, 2.62; II–4.10, 2.70; III–3.89, 2.41; IV–4.18, 2.61.

Trochantera: palp–length 1.95, width 2.10; I–2.95, 2.22; II–2.26, 2.27; III–2.15, 2.55; IV–2.60, 2.69.

Claws: paired as in female.

Male palp ( Figs 112–118): tibia mildly incrassate, tapering distally. Cymbium bipartite, tegulum large. Lanceolate embolus slightly tapers distally, little or no flaring, proximally emerges anterior to tegulum. Palpal tibia about three times length of embolus. Embolus terete (no keels) with distal “kiss curl” ( Fig. 116).

Distribution and natural history: To date, all known specimens (to the authors) of C. hoggi have come from the summit and south-western slopes of Fraser’s Hill, Selangor, West Malaysia, between elevations of 550-1150 meters, however, specimens of this species have recently been found at lower elevations on Fraser's Hill, nearer Kuala Kubu Bahru, Selangor, West Malaysia (J.-M. Verdez, pers. comm.). C. hoggi is a fossorial theraphosid that constructs its burrow retreat in steep sloped ground in shaded areas of montane tropical rainforest. Depending on the size of the specimen, burrows ranged between 20–60 cm in depth and terminate in an enlarged flask-like chamber. Burrow openings consist mainly of an elaborate silken collar that incorporates surrounding leaf litter and other organic debri. The soil is moist and clay-like. Males have been found between the months of January to March, during the monsoon season. Females have been found with egg sacs between March to September. Incubation times of selectively observed females were found to range from 24 to 28 weeks. Newly emerged C. hoggi instars have been observed in maternal burrows between September to late October. Ants were observed capturing and killing young instars as they dispersed from the maternal burrow (R. West and S. Hogg, pers. obs.).

Remarks: In 2000, R. West noted the presence of a new selenocosmiine species, located in Fraser’s Hill, Selangor, West Malaysia. Several specimens were gathered for scientific research under permit WL(AP/PP) 0227/00. A subsequent search for additional material via collection permit WL(AP/PP) 0244/01, again at Fraser’s Hill, yielded additional conspecifics.

Coremiocnemis jeremyhuffi sp. nov.

( Figs 119–151, 217)

Types: male holotype, 2 male paratypes, S83823 View Materials , S83821 View Materials , S83824 View Materials , respectively, Tapah (4° 06' N, 101° 25' E), Perak, West Malaysia, col. J.-M. Verdez, (date unknown), QM –examined; male paratype, ZRC ARA 743, Pahang, West Malaysia, col. F. N. Chasen, 193?, RMBR –examined; 2 female paratypes S88071 View Materials , S88072 View Materials , Fraser's Hill, Pahang, West Malaysia, col. S. Hogg, 22 Feb. 2009, QM –examined GoogleMaps .

Other material: 2 juveniles, S88069 View Materials , S88070 View Materials , Fraser's Hill , Pahang, West Malaysia, col. S. Hogg, 22 Feb. 2009, QM –examined; juvenile, S83755 View Materials , same data but, col. R. West, 7 April 2000, QM –examined; juvenile, S88021 View Materials , Tapah, Perak, West Malaysia, col. S. Yong, 24 Sept. 2008, QM –examined .

Etymology: A patronym in honour of Jeremy Huff who worked hard gathering specimens and data with the first author in Malaysia in 2000 and 2001.

Diagnosis: Differs from C. brachyramosa sp. nov. in its orange/brown color, increased bacillate count on prolateral maxillary surface and spermathecae with graduating sclerotization (not entire). Differs from C. gnathospina sp. nov. in smaller size and reduced interchelieral peg count (large pegs only). Differs from C. cunicularia , C. hoggi sp. nov., C. kotacana sp. nov., C. obscura sp. nov. and C. valida in having greatly reduced setation on metatarsi and tarsi IV, metatarsi IV retrolateral scopula <1/3 and lack of apically swollen spermathecal lobes.

Description: male holotype QM S83823 View Materials ( Fig. 120) with body length: 31.66.

Color (in life, Fig. 120): with creamy pinkish colored cephalothorax, dorsal coxae and trochantera. Femora jet black with paler colored setae, all other leg segments creamy/pink.

Carapace ( Fig. 121): length 11.20, and width 9.48 (width across anterior edge 5.80). Fovea 1.01, procurved, deep, under half width of OT ( Fig. 122).

Eyes ( Fig. 122): ocular tubercle; length 1.41, width 2.43. Anterior row transverse. Posterior row slightly recurved. Eyes: AME, ALE, PLE, PME.

Chelicerae ( Figs 128–130): length 7.05, width at base of each lobe 2.73. Ectal lyrate region a series of strikers (>60) in 7 horizontal rows ( Fig. 130). Strongest, longest strikers on lowest rows. Each striker highly needleform, largest with filiform ends. Teeth 9,>60 basomesals. Intercheliceral pegs (12 to 14 larger,>45 smaller) in tight cluster on basodorsal surface ( Fig. 129). Retrolateral face basomedially with singular spine, points distad.

Maxillae ( Figs 123, 126): prolaterally plano-convex, anterior lobe well pronounced, many cuspules (> 160) on inner basoventral surface. Maxillary lyra ( Figs 126, 127): large number of bacilliform rods (>450) in dense, ovoid patch on prolateral maxillary surface, lowest row composing up to 30 bacillae, longest rods medially in centre of lowest row. Rods paddle-shaped, short to medium shafts, largest lacking distal blades. At widest point, lyrate patch up to 16 rows deep, smallest rods dorsally. Posterior end of patch truncate, anterior end slightly pointed distad/rounded. Immediately above maxillary suture 20 elongated small spines on anterior margin of maxillae, rows unordered.

Labium ( Fig. 124): length 1.47, width 1.97. Many small cuspules (>650) along anterior 1/3 surface.

Sternum ( Figs 123, 125): length 5.01, width 4.45. Slightly wider posteriorly than anteriorly, with thin but even cover of short setae, interspersed with longer darker setae, most concentrated around posterior sternal borders. Posteriorly between left and right cox. IV, sternal border slightly acuminate, lateral sternal points lacking acumination. Sigilla ( Fig. 125): 3 pairs (not including labio-sternal sigilla), posterior medium, ovoid, 2.7 lengths apart, 1.7–1.9 of their length from sternal margin. Median pair 1.2 size of posterior, similar in form, 0.7–0.7 of their length from sternal margin. Anterior pair somewhat obscured, border sternal margin. Labio-sternal sigilla greatly obscured by setae, small.

Legs: formula; length IV, I, II, III; width III, I, IV, II. Leg RF ~ 99.97. Leg lengths (fem., pat., tib., met., tar., total): palp: 7.31, 4.18, 5.40, 0.00, 2.25, 18.04. I: 11.67, 5.74, 10.65, 8.93, 5.14, 42.13. II: 9.75, 4.55, 7.62, 7.10, 4.92, 33.94. III: 8.32, 4.29, 5.79, 6.96, 4.65, 30.01. IV: 11.45, 5.20, 9.65, 11.21, 4.63, 42.14. Leg mid– widths (fem., pat., tib., met., tar., total.): palp: 1.81, 1.75, 1.60, 0.00, 1.51, 6.67. I: 2.01, 2.33, 1.95, 1.29, 1.22, 8.80. II: 1.95, 2.05, 1.63, 1.07, 1.04, 7.74. III: 3.18, 2.23, 1.73, 1.07, 1.12, 9.33. IV: 2.19, 2.33, 1.48, 1.05, 1.01, 8.06.

Scopula: Met. and tar. I–III undivided, met. and tar. IV divided. Met. I scopula 7/8, met. II 5/6, met. III 2/ 3 and met. IV 1/4 ( Fig. 131).

Coxae: some shortened spiniform setae prolatero-dorsally, no thorns retrolaterally on I–IV. Coxae easily seen dorsally. Cox. I longest, about 1.2 times the length of II. IV widest, equal in length to III, basally rectangular with rounded corners. Coxae lacking ventral thorns (spiniform setae only) prolaterally on I–IV. I–III ventrally with many long thick blunt setae entirely and proximally, pallid. No short black setae. IV with mixture of long thick blunt setae proximally, pallid intermixed with shorter thin pallid setae. Ventral surface of coxae I–IV gently sloping anteriorly. Retrolateral setation: I–III with median narrow light brush, IV entirely pallid. I–IV retrolaterally lack ventral ledge. Ventral measurements for coxae (holotype only): palp–length 4.42, width 2.15; I–5.18, 2.72; II–4.35, 2.85; III–3.73, 3.01; IV–4.15, 3.01.

Trochantera: palp–length 2.06, width 2.00; I–3.06, 2.70; II–2.70, 2.63; III–2.35, 2.15; IV–2.48, 2.43.

Trichobothria: on all tarsi the basal filiform field slightly wider than clavate field, merges evenly. Clavates on tar. I in distal 2/3, long filiforms only in basal 1/2, shorter filiforms intermixed with clavates distally. Clavate extent on II–IV cf. I, in distal 2/3. Shorter filiforms for length. Short epitrichobothrial field on I shorter than clavates, uniform height for length.

Tarsal organ evident on legs I–II (under 40X).

Spines: met. I lacking spines, met. II with 1 short DV, met. III with 4 DV, 1 DL and met. IV with 4 DV, 2 DL.

Claws: paired claws on all legs and palpal claw unarmed. Reduced third claw on leg IV.

Abdomen: ovular, elongated, sandy brown, covered with mat of fine short dark setae, sparse cover of longer pale setae. Longer hairs more concentrated ventrally toward spinnerets, point distad. Slightly darker near booklungs.

Male palp ( Figs 132–136): tibia mildly incrassate, tapering distally. Cymbium bipartite, tegulum large. Lanceolate embolus slightly tapers distally with no flaring, proximally emerges anterior to tegulum. Palpal tibia ca. twice length of embolus. Embolus terete (no keels) with distal “kiss curl” ( Fig. 136).

Spinnerets: PMS: length 2.07, width (medially) 1.58. PLS: section lengths; basal 2.07, medial 1.58, apical 2.05. PLS section widths (medially); basal 1.04, medial 0.85, apical 0.64.

Female paratype (as in male, except) QM S88071 View Materials with body length: 39.78.

Color (in life, Fig. 119): dark slate femora, with paler distal leg segments, longer setae brightly colored, dulling in postultimate instars (R. West, pers. obs.).

Carapace ( Fig. 137): length 16.13, and width 12.46 (width across anterior edge 6.95). Fovea 2.56, procurved, equal width to OT ( Fig. 138).

Eyes ( Fig. 138): ocular tubercle; length 1.67, width 2.56. Anterior row transverse. Posterior row slightly recurved.

Chelicerae: length 6.59, width at base of each lobe 3.61. Teeth 11,>70 basomesals. Intercheliceral pegs (12 larger,>35 smaller) in tight cluster on basodorsal surface ( Fig. 144). Retrolateral face basomedially, 8 spiniform setae, point distad.

Maxillae ( Figs 139, 142): prolaterally plano-convex, anterior lobe well pronounced, many cuspules (> 200) on inner basoventral surface. Maxillary lyra ( Fig. 142): large number of bacilliform rods (>190) in dense, ovoid patch on prolateral maxillary surface, lowest row with>20 bacillae, longest rods medially within lowest row. Rods paddle-shaped, medium to long shafts, largest lack distal blades, however blackened tips evident. At widest point, lyrate patch>10 rows deep, smallest rods dorsally. Posterior end of patch truncate, anterior end slightly pointed distad/rounded. Dorsal to maxillary suture 33 stout small spines on anterior margin of maxillae, rows unordered.

Labium ( Fig. 140): length 2.34, width 2.86. Many small cuspules (>650) along anterior 1/2 surface.

Sternum ( Fig. 138): length 7.31, width 6.78. Sigilla ( Fig. 139): 3 pairs (not including labio-sternal sigilla), posterior medium, ovoid, length/width right: 0.89/0.41; left: 0.83/0.38, 1.05–1.25 from sternal margin. Median pair 1/2 size of posterior, similar in form, border sternal margin. Anterior pair obscured, border sternal margin. Labio-sternal sigilla greatly obscured by setae, small.

Legs: formula; length IV, I, II, III; width I, III, II, IV. Leg RF ~93.97. Leg lengths (fem., pat., tib., met., tar., total): palp: 8.89, 5.64, 6.51, 0.00, 5.93, 26.97. I: 11.96, 7.72, 10.07, 8.0, 5.91, 43.66. II: 11.39, 7.06, 8.63, 8.31, 5.87, 41.26. III: 9.71, 6.27, 6.27, 7.41, 4.76, 34.42. IV: 12.70, 6.82, 9.96, 12.02, 4.96, 46.46. Leg midwidths (fem., pat., tib., met., tar., total.): palp: 2.80, 2.59, 2.22, 0.00, 2.03, 9.64. I: 2.84, 3.0, 2.65, 2.29, 2.38, 13.16. II: 2.83, 2.62, 2.24, 1.99, 1.96, 11.64. III: 3.18, 2.81, 2.39, 1.69, 2.07, 12.14. IV: 2.86, 2.71, 2.15, 1.65, 1.63, 11.0.

Scopula: Met. and tar. I–III undivided, met. and tar. IV divided. All scopulae long, sparse. Metatarsus scopula (proximally): met. I entire, met. II 5/6, met. III 2/3, met. IV 1/5.

Coxae: Ventral measurements for coxae: palp–length 6.1, width 3.45; I–6.72, 3.15; II–4.12, 3.01; III–4.65, 3.05; IV–6.59, 4.69. Trochantera: palp–length 2.08, width 2.22; I–3.49, 3.28; II–2.89, 2.96; III–2.61, 2.93; IV–3.52, 3.32.

Tarsal organ evident on legs I–II (under 40X).

Claws: paired claws on all legs and palpal claw unarmed. Reduced third claw on leg IV.

Abdomen: ovular and elongated, sandy brown, covered with mat of fine short dark setae, with sparse cover of longer pale setae. Longer hairs more concentrated ventrally toward spinnerets, point distad. Slightly darker near booklungs.

Gentialia ( Fig. 141): epigastric fold, 4.35. Spermathecae paired, not fused, basally dividing (bilobular), with lateral lobes as long as medials. Lateral lobes robust, slightly swollen distally, medials lacking swollen tips. Sclerotization heaviest distally, gradually decreasing proximally. Right medial lobe in holotype with branching bud, extending laterally toward outer lateral bilobe. Bud emerges approx. 0.7 of total medial lobe length (proximally).

Spinnerets: PMS: length 1.91, width (medially) 0.43. PLS: section lengths; basal 2.56, medial 2.12, apical 3.01. PLS section widths (medially); basal 1.43, medial 1.25, apical 1.15.

Distribution and natural history: C. jeremyhuffi is known to range along the Titiwangsa Mountains east of Tapah, Perak, to the eastern slopes of Fraser’s Hill, Pahang, West Malaysia (see: Remarks). C. jeremyhuffi is a fossorial theraphosid that constructs its burrow retreat in almost vertical embankments in shaded areas of montane tropical rainforest between 250–1200 meters of elevation ( Fig. 148). Depending on the size of the specimen, burrows range between 20–40 cm in depth and terminate in an enlarged flask-like chamber. Burrow openings were found to be simple amongst very short vegetation and mosses ( Figs 149-151), with little to no traces of silk around the entrance (R. West, pers. obs.). Nothing else is known about the natural history of C. jeremyhuffi .

Remarks: The West Malaysian States of Selangor and Pahang divide the Fraser’s Hill region in a northwest–southeast direction, with the southwest slopes being in Selangor State and the northeast slopes being in Pahang State.

QM

Queensland Museum

RMBR

Raffles Museum of Biodiversity Research

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