Leiodes okawai, Nakane, 1963

Hoshina, Hideto, 2012, Review of the tribes Sogdini and Leiodini from Japan and North Chishima Islands. Part II. Genera Hydnobius and Leiodes (Coleoptera: Leiodidae), Acta Entomologica Musei Nationalis Pragae (suppl. 1) 52, pp. 1-168 : 94

Leiodes okawai species group

Species included. Leiodes kamezawai sp. nov., L. okawai Nakane, 1963 , L. yukihikoi sp. nov.

Diagnosis. Elytra unicolor and with sparsely arranged large punctures between rows of punctures ( Figs. 61D , 64D , 67D ); mesoventrite with distinct excavation between median carina and transverse carina ( Figs. 61E , 64E , 67E ); protarsi sexually dimorphic, tarsomeres 2–4 a little expanded in males ( Figs. 62A , 65A , 68A ); mesotibiae without sexual dimorphism, simply square at interoapical corner; male metafemur bearing prominent dorsal projection posteroapically, projection of female much smaller than in male ( Figs. 62G, 62H , 65G, 65H , 68G, 68H ); pubescence of metaventrite distinctly or indistinctly sexually dimorphic ( Figs. 61F, 61G , 64F, 64G , 67F, 67G ); aedeagus relatively small compared to body size, with one or a few small sclerites in inner sacs ( Figs. 63A , 66A , 69A ); male abdominal sternite 8 moderately or strongly curved ( Figs. 63C , 66D , 69C ); female abdominal sternite 8 with a spiculum ventrale at central point of anterior margin ( Figs. 63D , 66E , 69D ).

Differential diagnosis. Leiodes okawai species group can be separated from other species group by having the male metafemur bearing a prominent dorsal projection posteroapically and the aedeagus relatively small compared to the body size.

Figures

Fig. 61. Leiodes kamezawai sp. nov.A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.2 mm for E; IV: 1 mm for F and G.

Fig. 64. Leiodes okawai Nakane, 1963.A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 0.5 mm for C; III: 0.5 mm for E; IV: 1 mm for F and G.

Fig. 67. Leiodes yukihikoi sp. nov.A – body, dorsal view; B – ditto, lateral view; C – antenna; D – elytral punctures; E – mesoventrite, lateral view; F – male metaventrite; G – female metaventrite. Scale I: 1 mm for A and B; II: 1 mm for C; III: 0.5 mm for E; IV: 1 mm for F and G.

Fig. 62. Leiodes kamezawai sp. nov. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male hind leg, ventral view; D – female hind leg, ventral view; E – male protibia, dorsal view; F – female protibia, dorsal view; G – male metafemur, dorsal view; H – female metafemur, dorsal view. Scale I: 0.5 mm for A and B; II: 1 mm for C and D; III: 0.2 mm for E and F; IV: 0.2 mm for G and H.

Fig. 65. Leiodes okawai Nakane, 1963. A – male fore leg, ventral view; B – female fore leg, ventral view; C – male hind leg, ventral view; D – female hind leg, ventral view; E – male protibia, dorsal view; F – female protibia, dorsal view; G – male metafemur, dorsal view; H – female metafemur, dorsal view. Scale I: 0.5 mm for A and B; II: 1 mm for C and D; III: 0.2 mm for E and F; IV: 0.5 mm for G and H.

Fig. 68. Leiodes yukihikoi sp. nov.A – male fore leg, ventral view; B – female fore leg, ventral view; C – male hind leg, ventral view; D – female hind leg, ventral view; E – male protibia, dorsal view; F – female protibia, dorsal view; G – male metafemur, dorsal view; H – female metafemur, dorsal view. Scale I: 1 mm for A and B; II: 1 mm for C and D; III: 0.2 mm for E and F; IV: 0.2 mm for G and H.

Fig. 63. Leiodes kamezawai sp. nov.A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D; III: 0.1 mm for E.

Fig. 66. Leiodes okawai Nakane, 1963.A – aedeagus, dorsal view; B – ditto, lateral view; C – inner sac, dorsal view; D – male abdominal sternite 8; E – female abdominal sternite 8; F – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.1 mm for C; III 0.2 mm for D and E; IV: 0.1 mm for F.

Fig. 69. Leiodes yukihikoi sp. nov. A – aedeagus, dorsal view; B – ditto, lateral view; C – male abdominal sternite 8; D – female abdominal sternite 8; E – coxite and stylus. Scale I: 0.5 mm for A and B; II: 0.2 mm for C and D, and 0.1 mm for E.