Neocyrtopeltocoris triangularis (Knight) Wyniger & Schuh & Henry, 2023

Wyniger, Denise, Schuh, Randall T. & Henry, Thomas J., 2023, Revision of the North American Hallodapini (Insecta: Hemiptera: Heteroptera: Miridae: Phylinae), American Museum Novitates 2023 (3994), pp. 1-48 : 39-45

publication ID	https://doi.org/ 10.1206/3994.1
persistent identifier	https://treatment.plazi.org/id/03E8F838-FFDA-FB76-0950-E592FB0F7326
treatment provided by	Felipe
scientific name	Neocyrtopeltocoris triangularis (Knight)
status	comb. nov.

Treatment

Neocyrtopeltocoris triangularis (Knight) , new combination

Figures 7–9 View FIG View FIG View FIG , table 1

Sericophanes triangularis Knight. 1918: 81 View in CoL (original description).

Sericophanes albomaculatus Knight, 1930: 320 View in CoL (original description). New synonym.

Sericophanes fuscicornis Knight, 1968: 160 View in CoL (original description). New synonym.

Sericophanes nevadensis Knight, 1968: 160 View in CoL (original description). New synonym.

DIAGNOSIS: In addition to the characters outlined in the generic diagnosis, recognized by the usually orange general coloration (sometimes darker and more strongly brown) and the white marking on corium adjacent to apex of claval commissure. Most easily confused with Neocyrtopeltocoris froeschneri , n. sp. (fig. 7), in body form and color pattern, but that species smaller, with an overall castaneous or dark reddish-brown coloration, without the white marking on corium adjacent to apex of the claval commissure, and with a nonoverlapping distribution (fig. 9). Also potentially confused with Cyrtopeltocoris spp. , but members of that genus always with hemelytral fascia more or less parallel sided (figs. 1–3).

REDESCRIPTION: Male: Mean total length 3.66. Coloration (fig. 7): Orange to nearly brown, including appendages, tibiae more strongly reddish; corium with distinct white patch adjacent to apex of claval commissure. Surface and vestiture (fig. 7): Dorsum with long, erect, simple setae; antennae and legs with short, recumbent/appressed simple setae. Structure (fig. 7): As in generic description. genitalia (fig. 8): Endosoma long, filamentous, secondary gonopore preapical; phallotheca large with elongate apical portion, right angulate and bearing a dorsomedial spinelike projection; left paramere strongly elevated toward apex; right paramere short, rounded laterally, and tapering toward apex.

Female (fig. 7): Coloration of body and appendages similar to male. Structure as in generic description. GENITALIA: Not examined.

DISTRIBUTION (fig. 9): Range from southern California in the West to central Texas in the East, and from North Dakota in the north to Coahuila and Sonora, Mexico in the South .

DISCUSSION: We treat the nominal species Sericophanes albomaculatus Knight , S. fuscicornis Knight , and S. nevadensis Knight —all of which were described from single specimens—as junior synonyms of S. triangularis Knight based on examination of the type specimens and a substantial amount of additional material from across the range of what we construe as a single broadly distributed taxon.

Neocyrtopeltocoris albomaculatus Knight is more strongly castaneous and appears to have body proportions that are a bit shorter and broader than most specimens Knight examined in his description of N. triangularis .

A specimen bearing the locality label data listed by H.H. Knight for the holotype of Sericophanes fuscicornis Knight was found in the USNM collection but bore no identification label, a situation we attribute to a housekeeping problem by Knight or upon the transfer of his collection to the USNM. We have affixed an appropriate holotype label. We furthermore treat S. fuscicornis as a junior synonym of N. triangularis .

The apex of the endosoma in the four nominal species listed above shows some variation (see fig. 8), but it does not correlate with other observed variation, such as intensity of coloration or total body length. We have therefore recognized a single taxon having a very broad distribution.

We examined more specimens of this species than any other member of the North American Hallodapini . There is also a significant amount of plant data associated with N. triangularis , the preponderance from the genera Baccharis and Parthenium (Asteraceae) . These observations might suggest that the biology of this species is dissimilar to that of members of Cyrtopeltocoris spp. , most collections of which give little indication of phytophagy or host-plant preference, as we have argued above.

HOLOTYPE male of Sericophanes triangularis Knight : USA: New Mexico: Luna Co. : Deming, July 12, 1917, H.H. Knight [no PBI USI number] ( CUIC).

HOLOTYPE male of Sericophanes albomaculatus Knight : USA: Jeff Davis Co.: Ft. Davis Mts. , no date, O.C. Poling ( AMNH _ PBI 00286410 View Materials ) ( USNM).

HOLOTYPE male of Sericophanes fuscicornis Knight : USA: Arizona: Pima Co. : Empire Mountains, 31.88919 ° N 110.62568 ° W, 1524 m, 20 May 1928, A.A. Nichol, Baccharis sp. (Asteraceae) ( AMNH _PBI 00411127) ( USNM).

HOLOTYPE male of Sericophanes nevadensis Knight : USA: Nevada: Nye Co.: Mercury [M(T)], Aug. 23, 1965, Jose Merino [at laboratory light] ( AMNH _ PBI 00286411 View Materials ) ( USNM).

PARATYPES of S. triangularis : USA: Arizona: Cochise Co. : Bowie, 32.3264 ° N 109.487 ° W, 1147 m, 15 Jul 1917, H.H. Knight, 1♂ ( AMNH _PBI 00411145) ( USNM). Texas Pass, 32.06048 ° N 110.0791 ° W, 1524 m, 20 Jul 1917, H.H. Knight, 4♂ ( AMNH _PBI 00411141– AMNH _PBI 00411144) ( USNM). New Mexico: Luna Co. : Deming, 32.26861 ° N 107.75806 ° W, 12 Jul 1917, H.H. Knight, light trap, 13♂ ( AMNH _PBI 00411128– AMNH _PBI 00411140) ( USNM). South Dakota: Haakon Co.: Philip, 44.03943 ° N 101.6651 ° W, 660 m, 23 Jun 1923, H.C. Severin, Baccharis sp. (Asteraceae) , 1♂ ( AMNH _PBI 00411166) ( USNM).

OTHER SPECIMENS EXAMINED: MEXICO: Coahuila: Bilbao, 8 mi. N of Viesca, sand dunes, 25.46565 ° N 102.8 ° W, 30 May 1981, J. Doyen and J. Liebherr, Parthenium hysterophorus L. ( Asteraceae ), 1♂ (AMNH_PBI 00081263) (UCB). Sonora: 40 mi W of Moctezuma, 29.81666 ° N 110.33316 ° W, 27 Apr 1981, D.A. and J.T. Polhemus, 1♂ (AMNH_PBI 00392965) (JTP). USA: Arizona: Cochise Co. : Bowie, 32.3264 ° N 109.487 ° W, 1147 m, 15 Jul 1917, H.H. Knight, 1♂ (AMNH_PBI 00411152) (USNM). Chiricahua Mountains, Hidden Terrace, 4.5 mi SW of Portal, 31.8676 ° N 109.19499 ° W, 1646 m, 21 Jul 1982 – 25 Jul 1982, M.A. Cazier, 1♂ (AMNH_PBI 00392923) (AMNH). Douglas, 31.34444 ° N 109.54472 ° W, 29 Jul 1968, V.D. Roth, 3♂ (AMNH_PBI 00392931, AMNH_PBI 00392929, AMNH_PBI 00392930) (AMNH); 15 Aug 1968, V.D. Roth, 2♂ (AMNH_PBI 00392927, AMNH_PBI 00392928) (AMNH); 20 Aug 1969, V.D. Roth, 1♂ (AMNH_PBI 00392926) (AMNH). Huachuca Mountains, 5354 Ash Canyon Road, 0.5 mi W of Hwy 92, 31.38194 ° N 110.22444 ° W, 1554 m, 03 May 1992, N. McFarland, 1♂ (AMNH_ PBI 00411096) (USNM); 16 May 1992, N. McFarland, 1♂ (AMNH_PBI 00411097) (USNM); 21 Jun 1992, N. McFarland, 1♂ (AMNH_PBI 00411098) (USNM); 23 Jun 1992, N. McFarland, 3♂ (AMNH_ PBI 00411099–AMNH_PBI 00411101) (USNM); 01 Jul 1992, N. McFarland, 1♂ (AMNH_PBI 00411102) (USNM); 06 Jul 1992, N. McFarland, 2♂ (AMNH_PBI 00411103, AMNH_PBI 00411104) (USNM); 07 Jul 1992, N. McFarland, 1♂ (AMNH_PBI 00411106) (USNM); 09 Jul 1992, N. McFarland, 1♂ (AMNH_ PBI 00411107) (USNM); 17 Jul 1992, N. McFarland, 1♂ (AMNH_PBI 00411109) (USNM); 20 Jul 1992, N. McFarland, 1♂ (AMNH_PBI 00411110) (USNM); 16 Aug 1992, N. McFarland, 1♂ (AMNH_PBI 00411111) (USNM); 17 Aug 1992, N. McFarland, 1♂ (AMNH_PBI 00411112) (USNM); 18 Aug 1992, N. McFarland, 1♂ (AMNH_PBI 00411113) (USNM); 05 Sep 1992, N. McFarland, 1♂ (AMNH_PBI 00411115) (USNM); 12 Sep 1992, N. McFarland, 1♂ (AMNH_PBI 00411117) (USNM); 15 Sep 1992, N. McFarland, 2♂ (AMNH_PBI 00411116, AMNH_PBI 00411118) (USNM); 16 Sep 1992, N. McFarland, 2♂ (AMNH_PBI 00411114, AMNH_PBI 00411119) (USNM); 25 Sep 1992, N. McFarland, 1♂ (AMNH_PBI 00411120) (USNM). Portal, 31.91361 ° N 109.14083 ° W, 1500 m, 30 Aug 1976, J.D. Pinto, Parthenium hysterophorus L. ( Asteraceae ), 1♂ (AMNH_PBI 00083987) Parthenium hysterophorus L. ( Asteraceae ), 1♂ (AMNH_PBI 00083986) (UCR); 15 Jun 1980, R.T. Schuh, K. and R. Schmidt, light trap, 2♂ (AMNH_PBI 00175580, AMNH_PBI 00175581) (AMNH). Portal, 31.91333 ° N 109.14194 ° W, 11 Aug 1975, J.D. Pinto, 2♂ (AMNH_PBI 00083991, AMNH_PBI 00083992) (UCR). Coconino Co.: 3.5 mi S of Sedona on Rt 179, T17N R6E S30, 34.8255 ° N 111.769 ° W, 1280 m, 15 Jun 1983, R.T. Schuh and M.D. Schwartz, light trap, 1♂ (AMNH_PBI 00175558) (AMNH). Gila Co. : 8 mi SW jct Rts 87 and 188 (off Rt 87), Tonto National Forest, 33.55989 ° N 111.21341 ° W, 1219 m, 27 May 1983 – 28 May 1983, R. Schuh and G. Stonedahl, 1♂ (AMNH_PBI 00392925) (AMNH); 27 May 1983 – 28 May 1983, R.T. Schuh and G.M. Stonedahl, light trap, 2♂ (AMNH_PBI 00175556, AMNH_PBI 00175557) (AMNH). Mazatzal Wild[erness Reserve] along East Verde River, 34.2948 ° N 111.66321 ° W, 758 m, 30 Apr 1981, D.A. and J.T. Polhemus, 21♂ (AMNH_PBI 00392938–AMNH_PBI 00392947, AMNH_PBI 00392950–AMNH_ PBI 00392960) (JTP). Miami, 33.39667 ° N 110.87166 ° W, 22 Jul 1932, R.H. Beamer, 1♂ (AMNH_PBI 00391481) (KU). Graham Co.: Bonita, Post Creek Canyon, 32.6481 ° N 109.92617 ° W, 1692 m, 16 Jul 1917, H.H. Knight, 9♂ (AMNH_PBI 00411153–AMNH_PBI 00411161) (USNM). Fort Grant, Pinaleno Mts., 32.62285 ° N 109.94591 ° W, 2134 m, 15 Jul 1917 – 19 Jul 1917, Cornell Univ. Biological Expedition, 1♂ (AMNH_PBI 00076965) (CUIC). Pinaleno Mountains, Stockton Pass, 32.64083 ° N 109.84306 ° W, 1631 m, 01 Jun 1983 – 02 Jun 1983, R.T. Schuh, G.M. Stonedahl, Light Trap, 20♂ (AMNH_PBI 00175559– AMNH_PBI 00175572, AMNH_PBI 00175574–AMNH_PBI 00175579), 1♂ (AMNH_PBI 00175573) (AMNH). Maricopa Co. : E of Sunflower [CL 1634], 33.85356 ° N 111.44581 ° W, 1035 m, 10 Aug 1982, J.T. Polhemus, 1♂ (AMNH_PBI 00392961) (JTP). Reavis Ranch Trail, 33.49111 ° N 111.15472 ° W, 1097 m, 19 Apr 1982, D.A. and J.T. Polhemus, 1♂ (AMNH_PBI 00392963) (JTP). Salt River Canyon at Apache Lake, 33.55811 ° N 111.53153 ° W, 610 m, 28 Apr 1981, D.A. and J.T. Polhemus, 1♂ (AMNH_PBI 00411606) (JTP). Sunflower, 33.8642 ° N 111.46763 ° W, 10 Aug 1982, J.T. Polhemus, 1♂ (AMNH_PBI 00392964) (JTP). Navajo Co.: 10 mi W of Woodruff, 34.76667 ° N 110.21667 ° W, 1631 m, 20 Apr 1971, J. May, 5♂ (AMNH_PBI 00410932–AMNH_PBI 00410936) (USNM). Pima Co. : 30 mi E of Quijotoa, Lot 542 sub 336, 31.97644 ° N 111.73154 ° W, 853 m, 28 Aug 1927 – 29 Aug 1927, Cornell University, 6♂ (AMNH_PBI 00076959–AMNH_PBI 00076964) (CUIC). Baboquivari Mountains, 31.8 ° N 111.5 ° W, 07 Jul 1935, E.D. Ball, 1♀ (AMNH_PBI 00186383) (USNM). Black Dike Prospect, Sierrita, 31.93333 ° N 111.26666 ° W, 1143 m, 26 Jul 1916 – 29 Jul 1916, unknown collector, 1♂ (AMNH_PBI 00392922) (AMNH). Madera Canyon, Santa Rita Mountains, 31.72713 ° N 110.87364 ° W, 1580 m, 15 Aug 1964, E. Schlinger, M. Irwin & P. Rauch, Parthenium hysterophorus L. ( Asteraceae ), 1♂ (AMNH_PBI 00083988) (UCR). Madera Canyon, Santa Rita Mts., 31.73054 ° N 110.87947 ° W, 1463 m, 02 Oct 1963, V.L. Vesterby, 2♂ (AMNH_ PBI 00125665, AMNH_PBI 00125666) (UCD). Rincon Mountains, 32.20995 ° N 110.57526 ° W, 2286 m, 02 Sep 1928, A.A. Nichol, 1♂ (AMNH_PBI 00411162) (USNM). Sabino Canyon, 32.33012 ° N 110.79448 ° W, 901 m, 10 Aug 1953, G.D. Butler, 1♂ (AMNH_PBI 00186371) (USNM). Sabino Canyon, Santa Catalina Mountains, 32.192 ° N 110.4835 ° W, 843 m, 10 Aug 1953, G.D. Butler, 3♂ (AMNH_PBI 00410938–AMNH_PBI 00410940) (USNM). Santa Rita Exp[erimental]. Range, 31.88166 ° N 110.86569 ° W, 985 m, 14 Apr 1989, T.J. Henry and A.G. Wheeler, Jr., 5♂ (AMNH_PBI 00411121–AMNH_PBI 00411125) (USNM). Tucson, 32.22167 ° N 110.92583 ° W, 732 m, 19 Apr 1924, A.A. Nichol, 1♂ (AMNH_ PBI 00411149) (USNM); 07 Jun 1924, A.A. Nichol, 2♂ (AMNH_PBI 00411150, AMNH_PBI 00411151) (USNM). Pinal Co.: Sacaton, 33.07667 ° N 111.73861 ° W, 391 m, 06 May 1931, E.D. Ball, 1♀ (AMNH_PBI 00411163) (USNM). Sunspace Ranch near Oracle, 32.58592 ° N 110.83906 ° W, 1209 m, 30 Sep 1993, D.A. Polhemus, 1♂ (AMNH_PBI 00068139) (JTP). Santa Cruz Co.: Atascosa Mountains, Sycamore Canyon, 31.4384 ° N 111.1858 ° W, 1242 m, 03 Oct 1993, D.A. Polhemus, 1♂ (AMNH_PBI 00068143) (JTP). Calabasas Canyon, P.G., Atascosa Mountains, 31.45888 ° N 110.98583 ° W, 1049 m, 18 Apr 1981, D.A. Polhemus, 7♂ (AMNH_PBI 00392933–AMNH_PBI 00392937, AMNH_PBI 00411607, AMNH_PBI 00392924) (JTP), 1♂ (AMNH_PBI 00392932) (UCB), 1♂ (AMNH_PBI 00411126) (USNM). Santa Cruz River, 31.54063 ° N 111.02923 ° W, 1009 m, 06 Aug 1932, E.D. Ball, 1♂ (AMNH_PBI 00410941) (USNM). Yavapai Co.: 1 mi E of Yarnell, 34.22167 ° N 112.72918 ° W, 29 Apr 1981, D.A. and J.T. Polhemus, 2♂ (AMNH_PBI 00392948, AMNH_PBI 00392962) (JTP). Kirkland, Peeples Valley, Lot 542 sub 331, 34.3181 ° N 112.70757 ° W, 1372 m, 22 Aug 1927 – 24 Aug 1927, Cornell University, 6♂ (AMNH_PBI 00076944, AMNH_PBI 00076954–AMNH_PBI 00076958) (CUIC). Unknown co.: unknown locality, no date provided, 1♂ (AMNH_PBI 00410937) (USNM). California: San Bernardino Co.: 8 mi SE of Ivanpah, Lanfair Valley, 35.25878 ° N 115.20956 ° W, 25 May 1977, Saul Frommer, 1♂ (AMNH_PBI 00083993) (UCR). Colorado: Adams Co.: Denver, 39.91332 ° N 104.95324 ° W, 1640 m, 13 Jul 1909, W..J. Gerhard, Baccharis sp. (Asteraceae) , 1♂ (AMNH_PBI 00411167) (USNM). Elbert Co.: 7 mi E of Parker, 39.51501 ° N 104.64124 ° W, 1876 m, 18 Aug 1983, D.A. and J.T. Polhemus, 2♂ (AMNH_PBI 00392968, AMNH_PBI 00392969) (JTP). Otero Co.: La Junta, 37.985 ° N 103.54333 ° W, 04 Sep 1977, J.T. Polhemus, 3♂ (AMNH_PBI 00392966, AMNH_PBI 00392967, AMNH_PBI 00411608) (JTP). Unknown co.: Colo 2213, no date provided, 1♂ (AMNH_PBI 00186372) (USNM). Colorado, no date provided, C.F. Baker, Baccharis sp. (Asteraceae) , 1♂ (AMNH_PBI 00411168) (USNM). Idaho: Cassia Co. : 5 mi NE of Malta, 42.34507 ° N 113.31066 ° W, 1379 m, 16 Jul 1952, W.F. Barr, 1♂ (AMNH_PBI 00086780) (WFBM). Kansas: Decatur Co.: Decatur County, 39.8 ° N 100.46667 ° W, no date provided, F.X. Williams, 1♂ (AMNH_PBI 00391485) (KU). Nevada: Humboldt Co.: 40 mi SW of Denio, 41.47104 ° N 118.29838 ° W, 1257 m, 18 Jul 1927 – 19 Jul 1927, Cornell University, 9♂ (AMNH_PBI 00076945–AMNH_PBI 00076953) (CUIC). Nye Co.: 2.5 mi E of Gabbs on Route 844, 38.8 ° N 117.8 ° W, 1615 m, 01 Jul 1983, R.T. Schuh, M.D. Schwartz, 1♂ (AMNH_PBI 00175582), 14♂ (AMNH_PBI 00175587–AMNH_PBI 00175600), 3♂ (AMNH_PBI 00175584–AMNH_PBI 00175586) (AMNH). Beatty, 36.90855 ° N 116.7592 ° W, 1008 m, 23 Jun 1967, W. Gagne, Parthenium hysterophorus L. ( Asteraceae ), 2♂ (AMNH_PBI 00081256, AMNH_PBI 00081257) (UCB). Fairbanks Spring, Ash Meadows, 36.49023 ° N 116.34143 ° W, 689 m, 23 Jun 1951, Ted Frantz, Parthenium hysterophorus L. ( Asteraceae ), 1♂ (AMNH_PBI 00078576) (CAS). Mercury, M(T), “Lab,” 37.00766 ° N 116.11392 ° W, 1315 m, 15 Jul 1968, E. Beck, J. Merino, 1♂ (AMNH_PBI 00404969) (USNM).

Northumberland Canyon Rd., Toquima Ranges, T14N R44E Sec 31, 39.04652 ° N 117.00091 ° W, 1951 m, 28 Jun 1983, R.T. Schuh, M.D. Schwartz, Sarcobatus vermiculatus (Hook.) Torr. (Sarcobataceae) , det. NYBG staff, 1♂ (AMNH_PBI 00175583) (AMNH); 28 Jun 1983, R.T. Schuh and M.D. Schwartz, 2♂ (AMNH_PBI 00175601, AMNH_PBI 00175602) (AMNH). New Mexico: Bernalillo Co.: Tijeras, along I-40, 35.08083 ° N 106.3875 ° W, 1931 m, 01 Jun 1983, J.T. Polhemus, 8♂ (AMNH_PBI 00068138, AMNH_ PBI 00068140–AMNH_PBI 00068142, AMNH_PBI 00068144, AMNH_PBI 00068145, AMNH_PBI 00411604, AMNH_PBI 00411605) (JTP). Dona Ana Co.: Mesilla Park, 32.27 ° N 106.80028 ° W, 12 Jul 1917, H.H. Knight, 2♂ (AMNH_PBI 00411147, AMNH_PBI 00411148) (USNM). Organ, 32.42556 ° N 106.59611 ° W, 03 Jul 1940, D.E. Hardy, 2♂ (AMNH_PBI 00391484, AMNH_PBI 00391482) (KU). Hidalgo Co.: Double Adobe Ranch, Animas Mountains, 31.94889 ° N 108.80667 ° W, 1676 m, 15 Aug 1952, H.B. Leech and W.J. Gertsch, Parthenium hysterophorus L. ( Asteraceae ), 1♂ (AMNH_PBI 00078698) (CAS). Guadalupe Mountains, 31.36455 ° N 108.9881 ° W, 25 Aug 1971, E.E. Grissell & R.F. Denno, 2♂ (AMNH_PBI 00125671, AMNH_PBI 00125672) (UCD). Santa Fe Co.: 8 mi E of Los Alamos, 35.88806 ° N 106.16363 ° W, 04 Jul 1982, D.A. and J.T. Polhemus, 5♂ (AMNH_PBI 00392970–AMNH_PBI 00392974) (JTP). Sierra Co. : 25 mi W Tularosa, 33.07314 ° N 106.45097 ° W, 01 Jul 1940, D.E. Hardy, 1♂ (AMNH_ PBI 00391483), 1♀ (AMNH_PBI 00391394) (KU). North Dakota: Golden Valley Co.: Beach, 46.91667 ° N 104 ° W, 847 m, no date provided, C.N. Ainslie, Baccharis sp. (Asteraceae) , 1♂ (AMNH_PBI 00411164) (USNM). Oklahoma: Creek Co.: Stroud, 35.75525 ° N 96.64687 ° W, 262 m, 10 Aug 1941, R.H. Beamer, 1♂ (AMNH_PBI 00391392) (KU). Tillman Co.: Grandfield, 34.23067 ° N 98.68742 ° W, 05 Jul 1937, Standish-Kaiser, Baccharis sp. (Asteraceae) , 2♂ (AMNH_PBI 00411169, AMNH_PBI 00411170) (USNM). Woodward Co.: Alabaster Caverns State Park, 36.90194 ° N 99.14555 ° W, 570 m, 08 May 1974, G.F. and S. Hevel, 1♂ (AMNH_PBI 00271777) (USNM). South Dakota: Haakon Co.: Philip, 44.03943 ° N 101.6651 ° W, 660 m, 23 Jun 1923, H.C. Severin, 1♀ (AMNH_PBI 00404983) (CNC), Baccharis sp. (Asteraceae) , 1♂ (AMNH_PBI 00411165) (USNM). Texas: Bosque Co.: Texas [Norse], 31.75 ° N 97.66667 ° W, 262 m, between 1870 and 1880, [Belfrage Collection], 1♂ (AMNH_PBI 00411146) (USNM). Brewster Co.: Big Bend National Park, Panther Junction, 29.31748 ° N 103.21783 ° W, 1219 m, 31 Aug 1971, E.E. Grissell & R.F. Denno, 2♂ (AMNH_PBI 00125668, AMNH_PBI 00125669) (UCD). Randall Co.: Palo Duro State Park, near Amarillo, 34.93333 ° N 101.66222 ° W, 924 m, 27 Jun 1968, J.E. Slansky, 1♂ (AMNH_PBI 00125673) (UCD). Utah: San Juan Co.: Grand Flat near Collins Canyon, 37.42167 ° N 110.16056 ° W, 1707 m, 28 May 1984, D.A. and J.T. Polhemus, 3♂ (AMNH_PBI 00068149, AMNH_PBI 00068172, AMNH_PBI 00068181) (JTP). Sevier Co.: Richfield, 38.7725 ° N 112.08333 ° W, 15 Jul 1929, E.W. Davis, Baccharis sp. (Asteraceae) , 1♂ (AMNH_PBI 00411171) (USNM). Wyoming: Carbon Co.: Rawlins, 41.79111 ° N 107.23806 ° W, 05 Jul 1961, H. and A. Howden, 1♂ (AMNH_PBI 00404968) (CNC).