Centaurea amadanensis subsp. gymnoclada (Jaub. & Spach) Negaresh, 2018

Negaresh, Kazem & Rahiminejad, Mohammad Reza, 2018, A revision of Centaurea sect. Cynaroides (Asteraceae, Cardueae-Centaureinae), Phytotaxa 363 (1), pp. 448-450 : 448-450

3b. Centaurea amadanensis subsp. gymnoclada (Jaub. & Spach) Negaresh , stat. nov. ( Figs. 10 and 11 )

Phaeopappus gymnocladus Jaubert & Spach (1847: 15) . Phaeopappus decurrens var. gymnocladus (Jaub. & Spach) Bornmüller (1914: 413) . Tomanthea gymnoclada (Jaub. & Spach) Takhtajan (1939: 243) . Centaurea amadanensis var. gymnoclada (Jaub. & Spach) Wagenitz (1963: 191) . Type:— IRAN. In Mesopotamia (verisimiliter erronee), 1837, Aucher-Eloy 3197 (holotype P!: three sheets comprising one specimen).

Phaeopappus decurrens var. anoplos Bornmüller in Koeie (1945: 30). Type :— IRAN. Prov. Lurestan: 50 km E Khorram Abad, 1300 m, 31 May 1937, Koeie 807 (holotype C!).

Median phyllaries acute or with a short mucro, 0.5–3 mm long.

Taxonomic and distribution remarks: — Centaurea amadanensis subsp. gymnoclada is endemic to Isfahan, Lurestan , Kermanshah and Kurdistan Provinces, W and near Central part of Iran ( Fig. 5 ). This subspecies prefers elevations higher than those of C. amadanensis subsp. amadanensis .

Chromosome numbers: — Centaurea amadanensis subsp. gymnoclada is a diploid taxon with the chromosome number of 2 n = 2 x = 18 ( Fig. 9B and Table 1). Our results agree with Parishani et al. (2014) who reported the chromosome number 2 n = 2 x = 18 for C. amadanensis subsp. gymnoclada collected from Iran, Kurdistan, Saqqez to Marivan, after Soote village.

Selected specimens examined:— IRAN. Prov. Kurdistan: Chehel Chashmeh Mountains , 15 km E Marivan (Dezh Shahpur), 1400 m, Rechinger 42977 ( W!) ; 6 July 1971, Lamond 4568 ( E!) . Prov. Lurestan: Oshtoran Kuh, 20 km SE Dow Rud, near Saravaneh , 2000–2200 m, 6 June 1974, Rechinger 48112 ( E!) ; Dow Rud, Saravand bifurcate, 2127 m, 18 June 2011, Ranjbar & Negaresh 26965 ( BASU!) ; Ranjbar & Negaresh 29121 ( BASU!) . Prov. Kermanshah: Kamyaran-Ravansar road, 5 km after Alak village , 1428 m, 14 June 2012, Ranjbar & Negaresh 30545 ( BASU!) ; Ranjbar & Negaresh 30548 ( BASU!) ; Kamyaran to Ravansar, 5–15 km after Alak , 1418 m, 30 May 2012, Ranjbar & Negaresh 29687 ( BASU!) ; 5 km to Ravansar, Bave Geh village , 1418 m, 30 May 2012, Ranjbar & Negaresh 29909 ( BASU!) ; Ranjbar & Negaresh 29911 ( BASU!) ; Baghala Olya village , 1880 m, 19 May 2012, Ranjbar & Negaresh 29622 ( BASU!) ; Paveh toward Bayangan, 5 km to Bayangan , 1330 m, 29 May 2014, Negaresh & Asadbegy 128 (herb. Negaresh!); Javanroud toward Ravansar, 5 km to Ravansar, 1430 m, 27 May 2014, Negaresh & Asadbegy 152 (herb. Negaresh!). Prov. Isfahan: in mountains near Golpayegan, Strauss s.n. ( B!) .

Figures

FIGURE 5. Distribution of Centaurea amadanensis subsp. amadanensis (circles), C. amadanensis subsp. gymnoclada (squares), and C. amadanensis subsp. ranjbarii (triangles).

FIGURE 9. Somatic metaphases of Centaurea spp. A. Centaurea amadanensis subsp. amadanensis (2n = 18). B. C. amadanensis subsp. gymnoclada (2n = 18). C. C. nemecii (2n = 18). D. C. regia var. regia (2n = 18). E. C. regia var. purpurea. Scale A˗E = 2 μm.

FIGURE 10. Centaurea amadanensis Schultz Bipontinus subsp. gymnoclada (Jaub. & Spach) Negaresh (Aucher-Eloy 3197, P). A. inflorescence. B. basal leaves. C. phyllaries and achenses. D. lower part of habit. Scale A˗D = 2 cm.

FIGURE 11. Centaurea amadanensis Schultz Bipontinus subsp. gymnoclada (Jaub. & Spach) Negaresh (Ranjbar & Negaresh 30545, BASU). A. capitula. B. inflorescence. C. habit and habitat. D. leaves. Scale A = 1 cm, B = 3 cm, C = 10 cm, D = 6 cm.