Stenosfemuraia cumbre Huber

Huber, Bernhard A. & Villarreal, Osvaldo, 2020, On Venezuelan pholcid spiders (Araneae, Pholcidae), European Journal of Taxonomy 718, pp. 1-317: 277-281

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Stenosfemuraia cumbre Huber

sp. nov.

Stenosfemuraia cumbre Huber   sp. nov.

Figs 978–979 View Figs 972–979 , 988–998 View Figs 988–990 View Figs 991–998 , 1004–1007, 1064

Stenosfemuraia   sp. – Huber & Arias 2017: 497 View Cited Treatment , fig. 1.


Distinguished from congeners by combination of: armature of male chelicerae ( Fig. 994 View Figs 991–998 ; two pairs of frontal processes in distinctive arrangement); shape of procursus [ Figs 991–993 View Figs 991–998 ; with subdistal transparent process as in S. pilosa ( González-Sponga, 2005)   and S. exigua Huber   sp. nov., but slightly S-shaped in lateral view]; wide distal bulbal sclerite with subdistal side-branch ( Figs 995–997 View Figs 991–998 ); and epigynum (Fig. 1004–1005; relatively large anterior plate on large protruding whitish area, sclerotized part of posterior plate restricted to median half); from most known congeners (except S. cuadrata González-Sponga, 2005   ) also by long legs (male tibia 1>5.0, female tibia 1>3.0).


The species name refers to the type locality; noun in apposition.

Type material

VENEZUELA   – Aragua • ♂ holotype, ZFMK (Ar 22135), Henri Pittier National Park, forest near La Cumbre (10.3575° N, 67.5771° W), 1450 m a.s.l., 20 Feb. 2020 (B.A. Huber, O. Villarreal M.) GoogleMaps   .

Other material examined

VENEZUELA   – Aragua • 4 ♂♂, 3 ♀♀, ZFMK (Ar 22136), and 1 ♂, 1 juv. in pure ethanol, ZFMK (Ven20-161), same collection data as for holotype GoogleMaps   1 ♂, ZFMK (Ar 18256) Henri Pittier National Park, Rancho Grande (10.349° N, 67.684° W), ~ 1150 m a.s.l., in building, 12 Dec. 2002 (B.A. Huber) GoogleMaps   2 ♂♂, assigned tentatively (examined in 1999; not  reexamined), AMNH, Rancho Grande , 1200 m a.s.l., cloud forest, 1–10 Aug. 1987 (Bordan & Peck). – Carabobo   1 ♂, 1 ♀, MNHN (Ar 10556), with Eugène Simon’s labels “14651. Psil. San Esteban!” and “La Cumbre” [presumably La Cumbre between San Esteban and Maracay, 10.33° N, 68.00° W, i.e., not  the homonymous type locality!], Mar. 1888 (E. Simon) GoogleMaps   .


Male (holotype)

MEASUREMENTS. Total body length 3.2, carapace width 1.3. Distance PME–PME 100 µm; diameter PME 120 µm; distance PME–ALE 90 µm; diameter AME 35 µm; distance AME–AME 20 µm. Leg 1: 22.6 (5.6 +0.5 +5.6+8.9 +2.0), tibia 2: 3.5, tibia 3: 2.6, tibia 4: 3.3; tibia 1 L/d: 45. Femora 1–4 width (at half length): 0.23, 0.23, 0.24, 0.21.

COLOR (in ethanol). Carapace pale ochre-yellow with dark ochre median and lateral marginal bands, ocular area and clypeus also dark ochre; sternum ochre-yellow; legs ochre-yellow to light brown, with black rings on femora (subdistally), tibiae (proximally, subdistally) and metatarsi (indistinct proximal ring); abdomen pale greenish-gray, dorsally and laterally with dark bluish marks, ventrally with dark ochre transversal mark in gonopore area; book lung covers barely darkened.

BODY. Habitus as in Fig. 978 View Figs 972–979 . Ocular area moderately raised. Carapace high, with deep thoracic groove, posteriorly not  inflated. Clypeus unmodified. Sternum wider than long (0.86/0.60), unmodified. Abdomen globular, pointed at spinnerets.

CHELICERAE. As in Fig. 994 View Figs 991–998 , with two pairs of frontal processes in distinctive arrangement, without modified hairs.

PALPS. As in Figs 988–990 View Figs 988–990 , in general very similar to congeners (see, e.g., Huber & Arias 2017: figs 35– 36); coxa with retrolateral-ventral apophysis; trochanter barely modified; femur with retrolateral-ventral process proximally and large ventral apophysis distally (arrow in Fig. 993 View Figs 991–998 ); procursus ( Figs 991–993 View Figs 991–998 ) slightly S-shaped, with subdistal transparent process like S. pilosa   and S. exigua Huber   sp. nov.; genital bulb ( Figs 995–997 View Figs 991–998 ) with complex distal process, wide ventral sclerite with subdistal side-branch.

LEGS. Without spines, with curved hairs on legs 1 (femur, tibia, metatarsus) and 2 (tibia, metatarsus); vertical hairs in higher than usual density on femora (ventrally) and tibiae; retrolateral trichobothrium of tibia 1 at 5%; prolateral trichobothrium present on all leg tibiae; tarsus 1 with ~25 pseudosegments, mostly distinct.


In general similar to male ( Fig. 979 View Figs 972–979 ), carapace slightly less high, legs with few vertical hairs, without curved hairs. Tibia 1 in four females: 3.5, 3.7, 3.8, 3.8. Epigynum (Figs 1004–1005) with relatively large plate on large protruding whitish area; sclerotized part of posterior plate restricted to median half. Internal genitalia ( Figs 998 View Figs 991–998 , 1006–1007) with contiguous pore plates, median anterior receptacle, and pair of lateral anteriorly diverging sclerites (very similar S. exigua Huber   sp. nov.).


Known from three localities in the western Coastal Range of Venezuela   , in the states Aragua and Carabobo (Fig. 1064).

Figs 999–1007. Stenosfemuraia González-Sponga, 1998   ; epigyna, ventral and lateral views and cleared female genitalia, ventral and dorsal views. 999–1002. S. exigua Huber   sp. nov.; from La Guaira, El Limón (type locality; ZFMK Ar 22134). 1003. Stenosfemuraia   sp. ‘Ven18-10’ from Aragua, Colonia Tovar (ZFMK Ar 22127). 1004–1007. S. cumbre Huber   sp. nov.; from Aragua, Henri Pittier National Park (type locality; ZFMK Ar 22136).

Natural history

Most specimens were collected from small webs that were protruding from accumulations of leaves or in dense mosses and ferns up to one meter above the ground; some specimens were hiding under large leaves on the ground (e.g., dead palm leaves).


Germany, Bonn, Zoologische Forschungsinstitut und Museum "Alexander Koenig"


USA, New York, New York, American Museum of Natural History


France, Paris, Museum National d'Histoire Naturelle


Zoologisches Forschungsmuseum Alexander Koenig


American Museum of Natural History


Museum National d'Histoire Naturelle














Stenosfemuraia cumbre Huber

Huber, Bernhard A. & Villarreal, Osvaldo 2020