Groehnizomus oculiferans, DE FRANCESCO MAGNUSSEN & S.P. MÜLLER, 2022

Magnussen, Ilian De Francesco, Müller, Sandro P., Hammel, Jörg U., Kotthoff, Ulrich & Harms, Danilo, 2022, Diversity of schizomids (Arachnida: Schizomida) revealed by new fossil genera and species from mid-Cretaceous Burmese amber with implications for a Gondwanan origin of the Burma Terrane, Zoological Journal of the Linnean Society 196, pp. 792-844 : 810-818

publication ID	https://doi.org/ 10.1093/zoolinnean/zlac034
publication LSID	lsid:zoobank.org:pub:26533E9F-C53F-4051-B6B4-990E63C08C74
DOI	https://doi.org/10.5281/zenodo.7194390
persistent identifier	https://treatment.plazi.org/id/4291F40C-1BA1-4A9C-A668-0E1AA2E65B38
taxon LSID	lsid:zoobank.org:act:4291F40C-1BA1-4A9C-A668-0E1AA2E65B38
treatment provided by	Plazi
scientific name	Groehnizomus oculiferans
status	sp. nov.

Treatment

† GROEHNIZOMUS OCULIFERANS DE FRANCESCO

MAGNUSSEN & S.P. MÜLLER SP. NOV.

( FIGS 12–19 View Figure 12 View Figure 13 View Figure 14 View Figure 15 View Figure 16 View Figure 17 View Figure 18 View Figure 19 )

Zoobank registration: urn:lsid:zoobank.org:act:4291F40C-1BA1-4A9C-A668-0E1AA2E65B38

Type material: Holotype ( Figs 12A–D View Figure 12 , 13A–D View Figure 13 ), adult homeomorphic ♂, collection no. GPIH05049 , ex collection Carsten Gröhn (no. 11266); paratype ( Fig. 17A–D View Figure 17 ), adult heteromorphic ♂, collection no. GPIH05055 , ex collection Carsten Gröhn (no. 11210).

Diagnosis

† Groehnizomus oculiferans can be characterized by the following features: slender pedipalps; flagellum shape flat and subrhomboidal with a rounded tip.

Etymology

The specific name is derived from Latin oculus, eye and feree, to bear, referring to the presence of eyes in this specimen, which is a rare occurence in Recent schizomids. It is masculine in gender.

Description

Total length from the anterior process to base of the flagellum 2.56 mm.

Colour (in amber): Legs I–III, end of the propeltidium and metapeltidium yellowish brown, nearly transparent; anterior propeltidium, pedipalps, opisthosoma, legs IV and flagellum brownish red, original colouring equivocal.

Prosoma: Length 1.13 mm; propeltidium length 0.91 mm, width 0.35 mm; anterior process broad at the base with apical tip, length 0.08 mm; setation: one seta near the tip (visible only from directly above) followed by two setae, one next to the other. Propeltidium setation: only one more single median seta on the propeltidium between the anterior process and the eyes ( Fig. 14A, B View Figure 14 ). No other setae observed owing to taphonomic processes. Corneate eyes present, clearly elevated above the surface of the propeltidium. Mesopeltidia width 0.15 mm. Metapeltidium divided ( Fig. 14C, D View Figure 14 ), length 0.24 mm, width (from outer margin of left plate to outer margin of right plate) 0.52 mm. Anterior sternum with 10 + 2 sternapophysial setae, length 0.36 mm, width 0.24 mm. Posterior sternum with six setae, length 0.14 mm, width 0.15 mm.

Pedipalps: Without armature, except for mesal spur on trochater; trochanter produced distally ( Fig. 15A–F View Figure 15 ), setation: six large and strong ventral setae, four ventroectal, two mesal; femur: marginal longer than trochanter, setation: nine dorsal, a single seta, eight in a row, three ventroectal, four ventromesal; patella: nearly as long as the femur, setation: four dorsal, three mesal, two dorsoectal, four ectal, two ventroectal; tibia: shorter than patella, setation: five ventroectal, five ventromesal, four dorsal, five dorsoectal, three dorsomesal; tarsus: with asymmetrical tarsal spurs, spur length: 0.02 mm, setation: + three dorsal, three ventromesal, four dorsomesal, four dorsoectal, three ventroectal, two mesal; claw: bent downwards, about one-third of the length of tarsus.

Chelicerae: Setation of the mesal side of chelicerae not visible, except for G6-1.

Legs: Left leg I aborted at the trochanter, right one well preserved and complete, tarsus with seven segments, leg I ~1.54 times as long as the body; anterodorsal margin of femur of leg IV produced at an angle of ~90°, femur of leg IV is 3.4 times longer than deep, leg IV 1.18 times longer than the body, leg formula: 1423.

Opisthosoma: Length 1.43 mm, width 0.48 mm; tergites X–XII not elongated, tergite XII with a tiny posterodorsal process. Abdominal tergites setation: I: Dm, right one missing; II–VII: Dm; VIII: Dm, Dl1; IX: Dl1, Dl2; X: not visible; XI: Dl1; XII: Dm, Dl1, Dl2;

setation pattern: 2:2:2:2:2:2:2:4:4:?:2:6. Flagellum: dorsoventrally flattened, subrhomboidal in shape but with a rounded tip (holotype Fig. 16A–G View Figure 16 ; paratype Fig. 19A–D View Figure 19 ), length 0.26 mm, width 0.26 mm; stalk length 0.08 mm; setation: Dm1, Dl2 paired, Dm4, Dl3 paired, Vm1, Vm2 paired, Vm3 paired, Vl1 paired, Vm5, Vl2 paired.

Holotype further measurements: Pedipalp: trochanter 0.34mm; femur 0.36 mm; patella 0.37 mm; tibia 0.29 mm; tarsus 0.22 mm; claw 0.07 mm; total length 1.65 mm. Leg I: trochanter 0.18 mm; femur 1.01 mm; patella 1.24 mm; tibia 0.80 mm; metatarsus 0.31 mm; tarsus (tarsus segments) 0.41 mm (0.04 mm; 0.03 mm; 0.04 mm; 0.05 mm; 0.06 mm; 0.06 mm; 0.13 mm); total length 3.95 mm. Leg II: trochanter 0.11 mm; femur 0.69 mm; patella 0.31 mm; tibia 0.39 mm; metatarsus 0.40 mm; tarsus (tarsus segments) 0.30 mm (0.14 mm; 0.07 mm; 0.09 mm); claw 0.05 mm; total length 2.25 mm. Leg III: trochanter 0.15 mm; femur 0.58 mm; patella 0.22 mm; tibia 0.35 mm; metatarsus 0.38 mm; tarsus (tarsus segments) 0.30 mm (0.14 mm; 0.07 mm; 0.09 mm); claw 0.05 mm; total length 2.03 mm. Leg IV: trochanter 0.18 mm; femur 1.06 mm; patella 0.27 mm; tibia 0.61 mm; metatarsus 0.57 mm; tarsus (tarsus segments) 0.39 mm (0.16 mm; 0.08 mm; 0.15 mm); claw 0.07 mm; total length 3.01 mm.

Paratype further measurements: Total length from the anterior process to base of the flagellum: 2.56 mm. Prosoma length 1.18 mm; propeltidium length 0.77mm, width 0.35 mm. Pedipalp: trochanter 0.42 mm; femur 0.69 mm; patella 0.85 mm; tibia 0.41 mm; tarsus 0.25 mm; total length 2.62 mm. Leg I: trochanter 0.25 mm; femur> 1.27 mm; patella missing; tibia missing; metatarsus missing; tarsus missing; total length> 1.52 mm. Leg II: trochanter 0.13 mm; femur 0.65 mm; patella 0.38 mm; tibia 0.45 mm; metatarsus 0.31 mm; tarsus 0.34 mm; total length 2.26 mm. Leg III: trochanter 0.15 mm; femur 0.66 mm; patella 0.29 mm; tibia 0.37 mm; metatarsus 0.51 mm; tarsus 0.27 mm; total length 2.25 mm. Leg IV: trochanter 0.25 mm; femur 1.09 mm; patella 0.41 mm; tibia 0.68 mm; metatarsus 0.61 mm; tarsus 0.42 mm; total length 3.46 mm. Leg IV is 1.35 times longer than the body. Opistosoma: length 1.38 mm, width 0.51 mm. Flagellum: length 0.29 mm, width 0.23 mm; stalk length 0.08 mm.

Variations: Paratype with heteromorphic pedipalps, nearly as long as the body ( Fig. 18A, B View Figure 18 ), 0.62 times the body length in the holotype; femur of leg IV is 2.75

818 I. DE FRANCESCO MAGNUSSEN ET AL.

times longer than deep, 3.4 times longer than deep in the holotype.