Philonthus neuter, Mulsant and Rey, 1878

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium. I. Introduction, History, Biographical Sketches, and Omaliine Group, Bulletin of the American Museum of Natural History 2001 (265), pp. 1067-1806 : 1067-1806

neuter Mulsant and Rey, 1878 , see: baudii Fauvel, 1872 .

newelli Hatch, 1957: 105 ( Bledius ; subgenus Hesperophilus ; Type locality: Oregon: Lane Co.: Winchester Bay. Also cited from Washington).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1983: 77, 127 ( Bledius ; annularis group; characters; natural history; Oregon; Washington).

— Herman, 1986: 194, 319 ( Bledius ; annularis group; distribution; catalog).

- DISTRIBUTION: USA.

niger Cameron, 1945c: 707 ( Bledius ; subgenus Hesperophilus ; Type locality: South Africa: Simon’s Town).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1986: 252, 319 ( Bledius ; verres group; distribution; catalog).

- DISTRIBUTION: Angola, South Africa.

angolensis Cameron, 1950f: 115 ( Bledius ; subgenus Hesperophilus ; variety of niger ; Type locality: Angola: Dundo).

— Herman, 1986: 319 ( Bledius ; synonym of niger ).

nigricans Erichson, 1840 , see: dissimilis Erichson, 1840 .

nigriceps Notman, 1920 , see: politus Erichson, 1840 .

nigripennis Bernhauer, 1910 [Note: Nomen nudum], see: longulus Erichson, 1839 . nigripennis Gabriel, 1929 [Note: Nomen nudum], see: longulus Erichson, 1839 .

niloticus Erichson, 1840: 776 ( Bledius ; Type locality: Aegypto).

— Fauvel, 1875a: xiii [= 1875b: 215] ( Hesperophilus ; group 2).

— Mulsant and Rey, 1878: 654 ( Bledius ; subgenus Pucerus ).

— Fauvel, 1886: 25 ( Bledius ; notes; Algeria; Egypt).

— Fauvel, 1897d: 262 ( Bledius ; notes; distribution).

— Fauvel, 1902b: 71 ( Bledius ; cited as nitolicus; Algeria; Egypt; Syria; Senegal; Gabon; Ghana; Madagascar).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog).

— Eichelbaum, 1913: 117 ( Bledius ; Tanzania).

— Rambousek, 1922: 1006 ( Bledius ; British East Africa).

— Winkler, 1925: 347 ( Bledius ; subgenus Pucerus ).

— Scheerpeltz, 1933: 1119 ( Bledius ; catalog).

— C. Koch, 1934: 57, 60 ( Bledius ; subgenus Pucerus ; characters; notes).

— Cameron, 1950b: 213 ( Bledius ; Sudan).

— Cameron, 1952e: 828 ( Bledius ; Dahomey).

— Herman, 1970: 378 ( Bledius ).

— Frank, 1982: 11 ( Bledius ; list of parasites).

— Herman, 1986: 253, 319 ( Bledius ; verres group; distribution; catalog).

— Biswas and Sen Gupta, 1989: 16 ( Bledius ; subgenus Pucerus ; characters; India).

- DISTRIBUTION: Niger, Tunisia, Morocco, Senegal, Ivory Coast, Ghana, Benin, Gabon, Congo, Angola, South Africa, Tanzania, Madagascar, Kenya, Ethiopia, Sudan, Egypt, Israel, Lebanon, Syria, India.

nitidicollis Eppelsheim, 1885: 142 ( Bledius ; [preoccupied]; Type locality: West Africa).

— Fauvel, 1902b: 71 ( Bledius ; synonym of niloticus ).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; synonym of niloticus ).

— Herman, 1986: 319 ( Bledius ; synonym of niloticus ).

nitidiceps LeConte, 1877: 224 ( Bledius ; semiferrugineus group; Type locality: California:

Los Angeles; [Note: See lectotype designation by Herman, 1983]. Also cited from San Diego , California ).

— Casey, 1889a: 52 ( Bledius ; semiferrugineus group).

— Scheerpeltz, 1933: 1119 ( Bledius ; catalog).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1976: 150 ( Bledius ; armatus group; characters; natural history; California; Utah).

— Herman, 1983: 119, 138 ( Bledius ; lectotype designation: Los Angeles, California; Baja California; California) .

— Herman, 1986: 180, 319 ( Bledius ; armatus group; distribution; catalog).

- DISTRIBUTION: Mexico, USA.

nitidicollis Eppelsheim, 1885 , see: niloticus Erichson, 1840 .

nitidicollis LeConte, 1863a: 52 ( Bledius ; Type locality: New York; [Note: See lectotype designation by Herman, 1983]).

— LeConte, 1877: 223, 225 ( Bledius ; semiferrugineus group; characters).

— Casey, 1889a: 52 ( Bledius ; semiferrugineus group).

— Blatchley, 1910: 464, 465 ( Bledius ; characters).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog).

— Notman, 1920b: 697 ( Bledius ; characters).

— Scheerpeltz, 1933: 1119 ( Bledius ; catalog).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1972: 229 ( Bledius ; semiferrugineus group; characters; natural history; Manitoba; Arkansas; Illinois; Iowa; Kansas; Maryland; Missouri; Montana; Nebraska; Ohio).

— Frank, 1982: 11 ( Bledius ; list of parasites).

— Herman, 1983: 119, 138 ( Bledius ; lectotype designation; Arkansas; Illinois; Indiana; Kansas; Kentucky; Louisiana; Missouri; Montana; Nebraska; Ohio; South Dakota; Wisconsin) .

— Herman, 1986: 175, 319 ( Bledius ; semiferrugineus group; distribution; catalog).

— Downie and Arnett, 1996: 453 ( Bledius ; characters; USA).

- DISTRIBUTION: Canada, USA.

nodieri Cameron, 1939d: 205 ( Bledius ; subgenus Pucerus ; Type locality: Senegal: Badoumbe).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1986: 256, 319 ( Bledius ; verres group; distribution; catalog).

- DISTRIBUTION: Senegal.

normandi C. Koch, 1938 , see: tristis Aubé, 1843 .

norquincoensis Scheerpeltz, 1972: 91 ( Bledius ; subgenus Hesperophilus ; Type locality: Argentinien: Rio Negro, Norquinco, 900 m).

— Herman, 1986: 158, 319 ( Bledius ; emarginatus group; distribution; catalog).

- DISTRIBUTION: Argentina.

notialus Herman, 1976: 131 ( Bledius ; armatus group; Type locality: Oklahoma: Kay Co.:

1 mile east of Ponca City, Arkansas River. Also cited from Arkansas, Iowa, Kansas, Missouri, Oklahoma, Texas, Wisconsin).

— Herman, 1983: 138 ( Bledius ; Arkansas; Oklahoma).

— Herman, 1986: 180, 319 ( Bledius ; armatus group; distribution; catalog).

- DISTRIBUTION: USA.

nuchicornis Mulsant and Rey, 1861 , see: tricornis Herbst, 1784 .

obihiroensis Nakane, 1963b: 21 ( Bledius ; Type locality: Japan: Hokkaido: Tokachi, Obihiro). — Nakane, 1963a: 84 ( Bledius ; characters; habitus photograph; Japan).

— Herman, 1970: 378 ( Bledius ).

— Shibata, 1976: 137 ( Bledius ; Japan).

— Herman, 1986: 194, 320 ( Bledius ; annularis group; distribution; catalog).

- DISTRIBUTION: Japan.

obscurus Mulsant and Rey, 1870 , see: denticollis Fauvel, 1872 .

obscurus Motschulsky, 1860: 556 ( Bledius ; Type locality: fleuve Sélenga près de Kiahta en Mongolie).

— Hochhuth, 1862: 85 ( Bledius ; characters; Mongolia; Kirghiz Steppes). — Fauvel, 1875a: xii [= 1875b: 214] ( Bledius ; group 1).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog).

— Winkler, 1925: 346 ( Bledius ; subgenus Blediodes ).

— Herman, 1970: 378 ( Bledius ).

— Smetana, 1975c: 162 ( Bledius ; Mongolia).

— Herman, 1986: 194, 320 ( Bledius ; annularis group; distribution; catalog).

— Kashcheev, 1989: 294 ( Bledius ; subgenus Hesperophilus ; Mongolia).

- DISTRIBUTION: Mongolia, Kyrgyzstan.

obsoletus Fauvel, 1872: 207 [= 1873: 49] ( Bledius ; Type locality: France: Albi ;

Haut Pyrenees).

— Fauvel, 1875a: xiii [= 1875b: 215] ( Astycops , Bargus , Tadunus ; group 3).

— Mulsant and Rey, 1878: 525 ( Bledius ; subgenus Blediodes ; characters; France).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog).

— Winkler, 1925: 346 ( Bledius ; subgenus Blediodes ).

— Portevin, 1929: 412 ( Bledius ; characters; France).

— Scheerpeltz, 1933: 1119 ( Bledius ; catalog).

— L. Benick, 1943: 95 ( Bledius ; subgenus Blediodes ; characters).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1986: 194, 320 ( Bledius ; annularis group; distribution; catalog).

- DISTRIBUTION: France.

obtusicollis C. Koch, 1938 [Note: Nomen nudum], see: denticollis Fauvel, 1872 .

obtusus Sharp, 1889: 413 ( Bledius ; Type locality: Japan: Niigata).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog).

— Winkler, 1925: 346 ( Bledius ; subgenus Blediodes ).

— Scheerpeltz, 1933: 1119 ( Bledius ; catalog).

— Shibata, 1976: 137 ( Bledius ; Japan).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1986: 176, 320 ( Bledius ; semiferrugineus group; distribution; catalog).

— J. Li, 1993: 22 ( Bledius ; China).

- DISTRIBUTION: Japan, China.

occidentalis Bondroit, 1907a: 245 ( Bledius ; Type locality: Holland: Zwijn ).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog).

— Everts, 1922: 141 ( Bledius ; notes; distribution; characters).

— Winkler, 1925: 346 ( Bledius ; subgenus Blediodes ).

— Scheerpeltz, 1933: 1119 ( Bledius ; catalog).

— C. Koch, 1938b: 136 ( Bledius ; notes; Bosnia; Holland; Belgium; Germany; Croatia).

— West, 1942: 145 ( Bledius ; distribution; habitat).

— L. Benick, 1943: 95 ( Bledius ; subgenus Blediodes ; characters).

— Tottenham, 1949: 410 ( Bledius ; subgenus Hesperophilus ; Britain).

— Tottenham, 1954: 52 ( Bledius ; subgenus Hesperophilus ; characters; habitat; Britain).

— Horion, 1963: 277 ( Bledius ; subgenus Blediodes ; Belgium; Holland; France; Bosnia; Germany; Austria).

— Lohse, 1964: 97 ( Bledius ; subgenus Hesperophilus ; characters; central Europe).

— Scheerpeltz, 1968a: 33 ( Bledius ; subgenus Blediodes ; catalog).

— Herman, 1970: 378 ( Bledius ).

— Pope, 1977: 24 ( Bledius ; Britain).

— Herman, 1986: 176, 320 ( Bledius ; semiferrugineus group; distribution; catalog).

— Lucht, 1987: 94 ( Bledius ; checklist; Germany; Czechoslovakia; Austria; Benelux; Denmark). — Lohse and Lucht, 1989: 138 ( Bledius ; notes).

— Haghebaert, Bruge, and Drugmand, 1990: 38 ( Bledius ; Belgium).

— Boháč, 1993: 43 ( Bledius ; subgenus Hesperophilus ; Slovakia).

— Ciceroni and Zanetti, 1995: 13 ( Bledius ; Italy).

- DISTRIBUTION: Europe.

omega Herman, 1983: 45 , 127 ( Bledius ; annularis group; Type locality: Wisconsin: Ashland Co.: 15 miles SSE Ashland, Marengo River. Also cited from Alberta, Manitoba, Ontario, Quebec, Colorado, Michigan, Montana, New York, and North Dakota). — Herman, 1986: 184, 320 ( Bledius ; albonotatus group; distribution; catalog). — Downie and Arnett, 1996: 454 ( Bledius ; characters; USA). - DISTRIBUTION: Canada, USA.

opacicollis Eppelsheim, 1893: 59 ( Bledius ; Type locality: Ost-Sibirien, in der Nähe von Irkutsk, von Tunka bis Munku-Sardik im Sajan’schen Gebirge). — Poppius, 1909: 10 ( Bledius ; subgenus Blediodes ; Lena Valley, Siberia ). — Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog). — Winkler, 1925: 346 ( Bledius ; subgenus Blediodes ). — Scheerpeltz, 1933: 1119 ( Bledius ; catalog). — Herman, 1970: 378 ( Bledius ). — Smetana, 1975c: 162 ( Bledius ; Mongolia). — Herman, 1986: 194, 320 ( Bledius ; annularis group; distribution; catalog). — Kashcheev, 1989: 294 ( Bledius ; subgenus Hesperophilus ; Mongolia). — Ryabukhin, 1999: 38 ( Bledius ; subgenus Hesperophilus ; catalog; habitat; distribution; northeastern Russia). - DISTRIBUTION: Mongolia, Russia.

opacifrons LeConte, 1877: 224 ( Bledius ; semiferrugineus group; Type locality: San Diego, California; [Note: See lectotype designation by Herman, 1983]. Also cited from Los Angeles, California). — Casey, 1889a: 52 ( Bledius ; semiferrugineus group). — Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog). — Notman, 1920b: 697 ( Bledius ; characters). — Scheerpeltz, 1933: 1119 ( Bledius ; catalog). — Hatch, 1957: 99 ( Bledius ; subgenus Hesperophilus ; characters). — Herman, 1970: 378 ( Bledius ). — Herman, 1976: 153 ( Bledius ; armatus group; characters; natural history; California; Colorado; Oregon; Washington). — Herman, 1983: 119, 139 ( Bledius ; lectotype designation: San Diego, California; British Columbia; Manitoba; California; Montana; Oregon; Wyoming). — Herman, 1986: 180, 320 ( Bledius ; armatus group; distribution; catalog). - DISTRIBUTION: Canada, USA.

lectus Casey, 1889a: 49 ( Bledius ; armatus group; Type locality: California: Sonoma Co. ; [Note: See lectotype designation by Herman, 1983]). — Bernhauer and Schubert, 1911: 131 ( Bledius ; catalog). — Notman, 1920b: 696 ( Bledius ; characters). — Scheerpeltz, 1933: 1117 ( Bledius ; catalog). — Herman, 1970: 378 ( Bledius ). — Herman, 1976: 153 ( Bledius ; synonym of opacifrons ). — Herman, 1983: 119 ( Bledius ; lectotype designation). — Herman, 1986: 320 ( Bledius ; synonym of opacifrons ).

piceus Fall, 1910: 108 ( Bledius ; semiferrugineus group; Type locality: California: Pasadena; [Note: See lectotype designation by Herman, 1983]. Also cited from Oceanside, California). — Notman, 1920b: 697 ( Bledius ; characters). — Scheerpeltz, 1933: 1120 ( Bledius ; catalog). — Herman, 1970: 379 ( Bledius ). — Herman, 1976: 153 ( Bledius ; synonym of opacifrons ). — Herman, 1983: 120 ( Bledius ; lectotype designation: Pasadena, California). — Herman, 1986: 320 ( Bledius ; synonym of opacifrons ).

regularis Fall, 1910: 109 ( Bledius ; semiferrugineus group; Type locality: California: — Scheerpeltz, 1933: 1121 ( Bledius ; catalog).

— Hatch, 1957: 98 ( Bledius ; subgenus Hesperophilus ; characters; Washington; Idaho).

— Herman, 1970: 379 ( Bledius ).

— Herman, 1976: 153 ( Bledius ; synonym of opacifrons ).

— Herman, 1983: 120 ( Bledius ; lectotype designation).

— Herman, 1986: 320 ( Bledius ; synonym of opacifrons ).

specularis Fall, 1910: 109 ( Bledius ; semiferrugineus group; Type locality: California:

Point Reyes; [Note: See lectotype designation by Herman, 1983]).

— Notman, 1920b: 697 ( Bledius ; characters).

— Scheerpeltz, 1933: 1122 ( Bledius ; catalog).

— Herman, 1970: 379 ( Bledius ).

— Herman, 1976: 153 ( Bledius ; synonym of opacifrons ).

— Herman, 1983: 121 ( Bledius ; lectotype designation).

— Herman, 1986: 320 ( Bledius ; synonym of opacifrons ).

condonensis Hatch, 1957: 98 ( Bledius ; subgenus Hesperophilus ; Type locality: Oregon: Condon. Also cited from Washington and Idaho).

— Herman, 1970: 377 ( Bledius ).

— Herman, 1976: 153 ( Bledius ; synonym of opacifrons ).

— Herman, 1986: 320 ( Bledius ; synonym of opacifrons ).

opacinus Scheerpeltz, 1960b: 73 ( Bledius ; subgenus Hesperophilus ; Type locality: Afghanistan).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1986: 257, 320 ( Bledius ; verres group; distribution; catalog).

- DISTRIBUTION: Afghanistan.

opaculus LeConte, 1863a: 54 ( Bledius ; Type locality: Maine; [Note: See lectotype designation by Herman, 1983]).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog).

— LeConte, 1877: 232, 233 ( Bledius ; cordatus group; characters).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1976: 88 Bledius ; basalis group; characters; natural history; Newfoundland; Nova Scotia ; Prince Edward Island; Maryland; Massachusetts; New Jersey; North Carolina).

— Herman, 1983: 119, 139 ( Bledius ; lectotype designation; Maine; Massachusetts) .

— Herman, 1986: 202, 320 ( Bledius ; basalis group; distribution; catalog).

— Downie and Arnett, 1996: 453 ( Bledius ; characters; USA).

- DISTRIBUTION: Canada, USA.

opacus Block, 1799: 117 ( Staphylinus ; Type locality: Plauischer Grund bei Dresden).

— Erichson, 1839a: 581 ( Bledius ; characters; Germany).

— Heer, 1839: 210 ( Bledius ; characters; Switzerland).

— Erichson, 1840: 771 ( Bledius ; characters; Germany).

— Redtenbacher, 1849: 734 ( Bledius ; characters; Austria).

— Hardy, 1851: 49 ( Bledius ; notes; England).

— Fairmaire and Laboulbène, 1856: 603 ( Bledius ; characters; France).

— Redtenbacher, 1857: 229 ( Bledius ; characters; Austria).

— Kraatz, 1857d: 828 ( Bledius ; characters; Germany).

— Thomson, 1861: 119 ( Bledius ; characters; Scandinavia).

— Hochhuth, 1862: 86 ( Bledius ; notes).

— Schiødte, 1866: 148 ( Bargus ).

— Thomson, 1867: 290 ( Bledius ).

— Fauvel, 1872: 204 [= 1873: 46] ( Bledius ; characters; France; Scandinavia; Britain; Germany; Switzerland; Italy; Austria; Russia; Algeria; North America).

— Fauvel, 1873b: 128 [= 1873c: 21] ( Bledius ; Sardinia).

— Redtenbacher, 1874: 249 ( Bledius ; characters; Austria).

— Fauvel, 1875a: xiii [= 1875b: 215] ( Astycops , Bargus , Tadunus ; group 3).

— Mulsant and Rey, 1878: 599 ( Bledius ; subgenus Blediodes ; characters; France).

— Fauvel, 1886: 27 [= 1886a: 19] ( Bledius ; Algeria).

— Fowler, 1888: 370 ( Bledius ; characters; Britain).

— Seidlitz, 1891a: 381 ( Bledius ; subgenus Bledius ; characters).

— Ganglbauer, 1895: 619 ( Bledius ; subgenus Blediodes ; characters; Europe; Algeria;

North America).

— Fauvel, 1897d: 263 ( Bledius ; notes; distribution).

— Stierlin, 1900: 393 ( Bledius ; subgenus Bledius ; characters; Switzerland).

— Fauvel, 1902b: 72 ( Bledius ; Algeria).

— Sainte-Claire Deville, 1907: 91 ( Bledius ; characters; France).

— Reitter, 1909: 167 ( Bledius ; subgenus Blediodes ; characters).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog).

— Johansen, 1914: 539 ( Bledius ; subgenus Blediodes ; characters; Denmark).

— Everts, 1922: 139 ( Bledius ; notes; distribution).

— Winkler, 1925: 346 ( Bledius ; subgenus Blediodes ).

— Krogerus, 1925a ( Bledius ; characters; natural history; larva).

— Krogerus, 1925 ( Bledius ; natural history).

— Porta, 1926: 46 ( Bledius ; subgenus Blediodes ; characters).

— Portevin, 1929: 414 ( Bledius ; characters; France).

— Blair, 1931a: 1231 ( Bledius ; Scilly Islands).

— Brunier, 1931: 41–42 ( Bledius ; natural history).

— Scheerpeltz, 1933: 1119 ( Bledius ; catalog).

— Larsen, 1936 ( Bledius ; natural history).

— Breddin, 1936 ( Bledius ; natural history).

— Lengerken, 1939: 131–135 ( Bledius ; natural history).

— West, 1942: 143 ( Bledius ; distribution; habitat).

— L. Benick, 1943: 94 ( Bledius ; subgenus Blediodes ; characters).

— Tottenham, 1949: 411 ( Bledius ; subgenus Hesperophilus ; Britain).

— Larsen, 1953 ( Bledius ; natural history).

— Tottenham, 1954: 53 ( Bledius ; subgenus Hesperophilus ; characters; habitat; Britain).

— Steel, 1956b: 108 ( Bledius ; characters).

— Palm, 1961a: 52 ( Bledius ; characters; Sweden; Denmark; Norway; Finland).

— Horion, 1963: 269 ( Bledius ; subgenus Blediodes ; Algeria; Scotland, Scandinavia, and Finland to Polar region; Russia; central Europe; Spain; Italy; Sardinia; Dalmatia; Bosnia Hercegovina; Romania; Germany; Austria).

— Lohse, 1964: 95 ( Bledius ; subgenus Hesperophilus ; characters; central Europe).

— Pototskaia, 1967: 32 ( Bledius ; larval characters).

— Scheerpeltz, 1968a: 33 ( Bledius ; subgenus Blediodes ; catalog).

— Szujecki, 1968a: 716 ( Bledius ; Poland).

— Herman, 1970: 378 ( Bledius ).

— Smetana, 1975c: 161 ( Bledius ; notes; Mongolia).

— Pope, 1977: 24 ( Bledius ; Britain).

— Frank, 1982: 11 ( Bledius ; list of parasites).

— Lohse, 1982a: 116, 117 ( Bledius ; characters).

— Herman, 1986: 194, 320 ( Bledius ; annularis group; distribution; catalog).

— Lucht, 1987: 94 ( Bledius ; checklist; Germany; Poland; Czechoslovakia; Austria; Switzerland; France; Benelux; Denmark; Sweden).

— Kashcheev, 1989: 293 ( Bledius ; subgenus Hesperophilus ; Mongolia).

— Boháč, 1993: 43 ( Bledius ; subgenus Hesperophilus ; Slovakia; Czech Republic).

— Steidle and Dettner, 1995a ( Bledius ; defensive glands and secretions; phylogenetic discussion).

— Ciceroni and Zanetti, 1995: 13 ( Bledius ; Italy).

— Czarniawski and Staniec, 1997: 36 ( Bledius ; structure of opening of abdominal defensive reservoirs).

— Owen, 1997a: 148 ( Bledius ; collecting sites; Ireland).

— Staniec, 1999b: 47 ( Bledius ; third instar larval characters; Poland).

— Zerche, 2000: 52 ( Bledius ).

— Staniec, 2000: 35 ( Bledius ; characters of eggs and attachment disk; number of eggs in egg chamber; habitat of adults and eggs; Poland).

- DISTRIBUTION: Algeria, Europe, Russia, Iran, Mongolia, China.

divisus Marsham, 1802: 510 ( Staphylinus ; Type locality: Britain).

— Curtis, 1829: 29 ( Hesperophilus ).

— Stephens, 1829: 292 ( Hesperophilus ; Britain).

— Stephens, 1834: 310 ( Hesperophilus ; characters; Britain).

— Stephens, 1839: 417 ( Bledius ; characters; England).

— G. Waterhouse, 1858: 29 ( Bledius ; synonym of opacus ).

— Ganglbauer, 1895: 619 ( Bledius ; synonym of opacus ).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; synonym of opacus ).

— Steel, 1956b: 109 ( Bledius ; synonym of opacus ).

— Herman, 1986: 321 ( Bledius ; synonym of opacus ).

castaneipennis Mannerheim, 1830: 45 ( Bledius ; Type locality: Suecia; Petropoli; Finlandia). — Mannerheim, 1831: 459 ( Bledius ; Finland; Sweden; Russia).

— Stephens, 1829: 292 ( Bledius ; nomen nudum).

— Gistel, 1834: 9 ( Dicarenus ).

— C. Sahlberg, 1831: 408 ( Oxytelus ; characters; Finland).

— Lacordaire, 1835: 457 ( Bledius ).

— Erichson, 1839a: 581 ( Bledius ; synonym of opacus ).

— Redtenbacher, 1849: 734 ( Bledius ; synonym of opacus ).

— Fairmaire and Laboulbène, 1856: 603 ( Bledius ; characters; France).

— Motschulsky, 1858d: 71 ( Bledius ; synonym of opacus ).

— Kraatz, 1857d: 828 ( Bledius ; synonym of opacus ).

— Mulsant and Rey, 1878: 599 ( Bledius ; synonym of opacus ).

— Seidlitz, 1891a: 381 ( Bledius ; synonym of opacus ).

— Ganglbauer, 1895: 619 ( Bledius ; synonym of opacus ).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; synonym of opacus ).

— Steel, 1956b: 109 ( Bledius ; synonym of opacus ).

— Herman, 1986: 173, 301 ( Bledius ; semiferrugineus group; distribution; catalog).

haemopterus Stephens, 1832: 310 ( Hesperophilus ; Type locality: Britain: Suffolk; Norfolk; Yorkshire ).

— Curtis, 1829: 29 ( Hesperophilus , nomen nudum).

— Stephens, 1829: 292 ( Bledius ; nomen nudum).

— Stephens, 1834: 310 ( Hesperophilus ; characters; Britain).

— Stephens, 1839: 417 ( Bledius ; characters; England).

— Hardy, 1851: 49 ( Bledius ; synonym of opacus ).

— G. Waterhouse, 1858: 29 ( Bledius ; synonym of opacus ).

— Ganglbauer, 1895: 619 ( Bledius ; synonym of opacus ).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; synonym of opacus ).

— Steel, 1956b: 109 ( Bledius ; synonym of opacus ).

— Herman, 1986: 321 ( Bledius ; synonym of opacus ).

extensus Motschulsky, 1860: 555 ( Bledius ; Type locality: Daourie).

— Fauvel, 1872: 204 [= 1873: 46] ( Bledius ; synonym of opacus ).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; variety of opacus ).

— Winkler, 1925: 346 ( Bledius ; variety of opacus ).

— Steel, 1956b: 109 ( Bledius ; synonym of opacus ).

— Herman, 1986: 321 ( Bledius ; synonym of opacus ).

subsinuatus Mulsant and Rey, 1878: 602 ( Bledius ; subgenus Blediodes ; Type locality: Grande Chartreuse ).

— Fauvel, 1895a: 118 ( Bledius ; synonym of denticollis ).

— Bernhauer and Schubert, 1911: 128 ( Bledius ; synonym of denticollis ).

— C. Koch, 1938b: 132 ( Bledius ; variety of opacus ; notes).

— Scheerpeltz, 1968a: 32 ( Bledius ; synonym of denticollis ).

— Lohse, 1982a: 116, 117 ( Bledius ; synonym of opacus ).

— Herman, 1986: 321 ( Bledius ; synonym of opacus ).

— Lucht, 1987: 94 ( Bledius ; checklist; Germany; Poland).

— Lohse and Lucht, 1989: 136 ( Bledius ; synonym of opacus ).

— Boháč, 1993: 43 ( Bledius ; synonym of opacus ).

sinuatocollis Gerhardt, 1899: 218 ( Bledius ; variety of opacus ; Type locality: Schlesien: Neisse).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; variety of opacus ).

— C. Koch, 1938b: 132 ( Bledius ; synonym of subsinuatus ).

— Steel, 1956b: 109 ( Bledius ; synonym of opacus variety subsinuatus ).

— Lohse, 1964: 94 ( Bledius ; synonym of subsinuatus ).

— Herman, 1986: 321 ( Bledius ; synonym of opacus ).

minor Krogerus, 1925: 117 ( Bledius ; variety of opacus ; [preoccupied]; Type locality: Ortschaft Malm im Kirchspiel Helsinge).

— Steel, 1956b: 109 ( Bledius ; synonym of variety krogeri ).

— Herman, 1986: 321 ( Bledius ; synonym of opacus ).

krogeri Scheerpeltz, 1933: 1119 ( Bledius ; replacement name for minor Krogerus ).

— Steel, 1956b: 109 ( Bledius ; appears to be species other than opacus ).

— Herman, 1986: 321 ( Bledius ; synonym of opacus ).

oregonensis Hatch, 1957 , see: suturalis LeConte, 1863 .

orientalis Bernhauer and Schubert, 1911: 133 ( Bledius ; replacement name for lividipes Fairmaire ).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1986: 218, 321 ( Bledius ; punctatissimus group; distribution; catalog).

- DISTRIBUTION: Djibouti.

lividipes Fairmaire, 1892: 90 ( Bledius ; [preoccupied]; Type locality: Obock).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; synonym of orientalis ).

— Herman, 1986: 321 ( Bledius ; synonym of orientalis ).

orientalis Roubal, 1934 , see: roubali Horion, 1963 .

orion Normand, 1939: 485 ( Bledius ; Type locality: Tunisia: Fernana ; Souk-el-Arba).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1986: 176, 321 ( Bledius ; semiferrugineus group; distribution; catalog).

- DISTRIBUTION: Tunisia.

ornatus LeConte, 1863 , see: albonotatus Mäklin, 1853 .

orphanus Sharp, 1874: 91 ( Bledius ; Type locality: Japan: Kobé ).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog).

— Bernhauer, 1922c: 224 ( Bledius ; characters; Formosa).

— Winkler, 1925: 347 ( Bledius ; subgenus Hesperophilus ).

— Scheerpeltz, 1933: 1119 ( Bledius ; catalog).

— Nakane, 1963a: 84 ( Bledius ; cited as orphinus; characters; habitus photograph; Japan).

— Herman, 1970: 378 ( Bledius ).

— Shibata, 1973c: 34 ( Bledius ; subgenus Hesperophilus ; Taiwan).

— Shibata, 1976: 138 ( Bledius ; Japan).

— Herman, 1986: 257, 321 ( Bledius ; verres group; distribution; catalog).

— J. Li, 1993: 21 ( Bledius ; China).

- DISTRIBUTION: China, Japan, Taiwan.

osborni † Scudder, 1900: 72 ( Bledius ; fossil; Type locality: Florissant , Colorado) .

— Handlirsch, 1907: 729 ( Bledius ; catalog; Miocene; Colorado).

— Wickham, 1912: 12 ( Bledius ; Colorado).

— Herman, 1986: 68, 69, 321 ( Bledius ; catalog).

osiris Normand, 1935: 361 ( Bledius ; Type locality: Tunisia: Le Kef).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1986: 195, 321 ( Bledius ; annularis group; distribution; catalog).

— Ciceroni and Zanetti, 1995: 13, 59 ( Bledius ; Italy?).

- DISTRIBUTION: Tunisia.

burlinii C. Koch, 1938b: 140 ( Bledius ; subgenus Blediodes ; subspecies of osiris ; Type locality: Italy: Lovadina near Treviso).

— Porta, 1949: 140 ( Bledius ; characters; Italy).

— Herman, 1986: 321 ( Bledius ; subspecies of osiris ).

— Ciceroni and Zanetti, 1995: 13 ( Bledius ; subspecies of osiris ; possible synonym of erraticus ; Italy).

ouzbekiscus Coiffait, 1970d: 145 ( Bledius ; subgenus Elbidus ; Type locality: Ouzbekistan: Urgench).

— Herman, 1986: 151, 321 ( Bledius ; kochi group; distribution; catalog).

— Kashcheev, 1991: 114 [= 1992: 167] ( Bledius ; subgenus Elbidus ; characters; Uzbekistan).

- DISTRIBUTION: Uzbekistan.

ovensensis Blackburn, 1891: 76 ( Bledius ; Type locality: Australia: Victoria: Ovens River). — Bernhauer and Schubert, 1911: 133 ( Bledius ; cited as ovensis; catalog).

— Scheerpeltz, 1933: 1119 ( Bledius ; cited as ovensis; catalog).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1986: 158, 321 ( Bledius ; emarginatus group; distribution; catalog).

- DISTRIBUTION: Australia.

palliatus Fauvel, 1895: 203 ( Bledius ; Type locality: Bírmanie: Myeenkyan ).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog).

— Bernhauer, 1922a: 149 ( Bledius ; Cambodia).

— Cameron, 1930: 278 ( Bledius ; subgenus Pucerus ; characters; Burma).

— Scheerpeltz, 1933: 1119 ( Bledius ; catalog).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1986: 258, 321 ( Bledius ; verres group; distribution; catalog).

- DISTRIBUTION: Myanmar, Cambodia.

pallidipennis Bernhauer, 1927e: 234 ( Bledius ; Type locality: Argentinien: Chaunar Region [Note: In a discussion of Neobisnius parcepunctatus, Frank, 1981: 12 , 13 thought this locality probably referred to Chañar Region, La Rioja]).

— Scheerpeltz, 1933: 1119 ( Bledius ; catalog).

— Herman, 1970: 378 ( Bledius ).

— Herman, 1986: 158, 321 ( Bledius ; emarginatus group; distribution; catalog).

- DISTRIBUTION: Argentina.

pallipennis Say, 1823: 155 ( Oxytelus ; Type locality: Missouri River below confluence of the Platte River).

— Erichson, 1840: 779 ( Bledius ; characters).

— LeConte, 1877: 218 ( Bledius ; mandibularis group; characters; Nebraska; Illinois).

— Blatchley, 1910: 464 ( Bledius ; characters; Indiana).

— Bernhauer and Schubert, 1911: 133 ( Bledius ; catalog).

— Herman, 1970: 378 ( Bledius ).

— Smith and Hein, 1971 ( Bledius ; natural history; Nebraska).

— Herman, 1972: 173 ( Bledius ; mandibularis group; characters; natural history; Arkansas; Illinois; Iowa; Kansas; Louisiana; Mississippi; Missouri; Nebraska; New Mexico;

North Dakota; Ohio; Oklahoma; South Dakota; Texas).

— Herman, 1983: 139 ( Bledius ; Kansas; Mississippi; Missouri; Nebraska; Texas).

— Herman, 1986: 112, 321 ( Bledius ; mandibularis group; distribution; catalog).

— Downie and Arnett, 1996: 452 ( Bledius ; characters; USA).

- DISTRIBUTION: USA.