Achilia cunniceps, Sabella & Cuccodoro & Kurbatov, 2019

Sabella, Giorgio, Cuccodoro, Giulio & Kurbatov, Sergey A., 2019, A revision of the Chilean Brachyglutini - Part 4. Revision of Achilia Reitter, 1890: A. puncticeps and A. approximans species groups, with description of seven new species (Coleoptera: Staphylinidae: Pselaphinae), Revue suisse de Zoologie 126 (1), pp. 127-145 : 137

publication ID	https://doi.org/ 10.5281/zenodo.2619528
DOI	https://doi.org/10.5281/zenodo.3706247
persistent identifier	https://treatment.plazi.org/id/03E587B3-0D30-3305-EB54-E0C06D6AFB56
treatment provided by	Carolina
scientific name	Achilia cunniceps
status	sp. nov.

Treatment

Achilia cunniceps View in CoL n. sp.

Figs 7 View Figs 7-10 , 11 View Figs 11-14 , 23 View Figs 19-28 , 35, 37, 39 View Figs 35-40 , 57 View Fig

Holotype: MHNG (# MHNG-ENTO-13837); 1 ♂; SOUTHERN CHILI: Región Los Lagos: Osorno prov.: Puyehue National Park, Antillanca road; 500-1000 m; 18-20.XII.1984; S. & J. Peck; car netting.

Paratypes (3): MHNG (# MHNG-ENTO-13838 & 13839); 2 ♂; SOUTHERN CHILI: Región Los Lagos: Osorno prov. : same data as holotype . – Chiloé prov.: FMNH (FMHD # 2002-77); 1 ♂; road to Miraflores, about 0.6 km W road 5; 42° 46.74’S 73° 47.70’W; 130 m; 12.XII.2002; A. Newton & M. Thayer 1063; secondary Valdivian forest with few conifers, Berlese, leaf & log litter GoogleMaps .

Description: Body 1.35-1.40 mm long, entirely dark with reddish brown elytra, antennae, palpi, and legs. Head with eyes moderately protruding, about long as convex temples. Pronotum slightly wider than head; median antebasal fovea smaller than lateral ones. First abdominal tergite with basal striae extending to about one-third of paratergal length, and separated at base by about one-third of tergal width.

Male: Head as in Figs 35, 37, 39 View Figs 35-40 , with anterior part flattened; vertexal sulcus indistinct; posterior part conspicuously raised, strongly convex in lateral view ( Fig. 37 View Figs 35-40 ), this hump anteriorly flattened, densely punctate, pubescent, and sulcate medially. Antennae ( Fig. 11 View Figs 11-14 ) with scape longer than wide and pedicel only slightly longer than wide; antennomeres III-IV and VII slightly wider than long; antennomeres V-VI longer than wide, antennomere VIII strongly transverse with mesal margin protruding at middle; antennomeres IX and X strongly transverse with protruding mesal margins; antennomere XI very elongate and distinctly longer than VI-X combined, its surfaces with tubercles. Metaventrite raised at middle, this area distally pubescent and densely punctate, divided by very wide median sulcus. Legs with trochanters very elongate; surface of mesotrochanters bearing numerous long bristles; ventral margin of mesotrochanters projecting posteriorly in a spine ( Fig. 23 View Figs 19-28 ); profemora and mesofemora slightly thickened; protibiae with short subapical spur on medial margin, distal half slightly sinuate and bearing long and thick bristles; distal half of metatibiae slightly sinuate. Abdominal tergites and ventrites unmodified. Aedeagus ( Fig. 7 View Figs 7-10 , dorsal longitudinal struts not shown) 0.26- 0.27 mm long; medial sclerites with 5-6 apical spines, associated on each side with three pointed sclerites, the first with two tips.

Female: Unknown.

Collecting data: Collected in December, at elevations ranging from 130 to 1000 m, by car netting, and by sifting in Valdivian rainforests.

Distribution: Achilia cunniceps n. sp. is known from Southern Chile ( Fig. 57 View Fig : blue triangles) only in Región Los Lagos (Chiloé and Osorno provinces).

Comments: Achilia cunniceps n. sp. is easily distinguished from the other members of the genus by the very peculiar morphology of the male head ( Figs 35, 37, 39 View Figs 35-40 ), features of the antennae ( Fig. 11 View Figs 11-14 ), and the copulatory pieces of the aedeagus ( Fig. 7 View Figs 7-10 ).