Abida secale secale ( Draparnaud, 1801 )

Kokshoorn, Bas & Gittenberger, Edmund, 2010, Chondrinidae taxonomy revisited: New synonymies, new taxa, and a checklist of species and subspecies (Mollusca: Gastropoda: Pulmonata) 2539, Zootaxa 2539 (1), pp. 1-62 : 13

Abida secale secale ( Draparnaud, 1801) (pl. 1 figs A–J)

Pupa secale Draparnaud, 1801: 59 .

Type locality: France.

Lectotype: Museum of Natural History , Vienna 77709.

Distribution.— England, Belgium, southern Germany, Slovakia, northwestern Italy, France and northeastern Spain. See Kerney, Cameron & Jungbluth (1983: 332, fig. 86).

Description (after Gittenberger 1973).—Shell more or less slender, cylindro-conical, with 8¼–9⅝ weakly to moderately inflated whorls, sculptured with irregularly placed, weak to rather prominent axial ribs. Body whorl obliquely flattened, not or only slightly narrowed, with an indistinct keel and an indistinct indentation at the position of both the infrapalatalis and the palatalis superior. The palatal side of the aperture slightly leaning forward (in lateral view). Umbilicus open, partly obscured by the last part of the body whorl. Shell height, 6.0– 9.5 mm; width, 2.0– 2.9 mm.

The parietal edges of the aperture are connected by a faint to barely visible callus. Angularis mostly connected with spiralis. Subangularis partly fused with angularis. Parietalis present. Columellaris reaching further than the columella in front, but not to the edge of the aperture. Infracolumellaris as prominent as the columellaris or somewhat smaller. A third, much less developed columellar fold, the basalis, is often present. Infrapalatalis, palatalis inferior and palatalis superior are hardly to clearly indented or interrupted, reaching in front to the edge of the aperture. At the anterodorsal palatal centre a suprapalatalis and a suturalis are present.

Genetic barcode.—GenBank accession numbers EU395399 – EU395415 , EU395427 .

Notes.— Abida secale secale is somewhat variable in size across its range (see pl. 1 figs G, H). However, the apertural dentition is very uniform. This also accounts for samples from high altitudes in the Alps (i.e. pl. 1 fig. I). This is strikingly different from the situation in the Pyrenees, where shell morphology varies conspicuously with the altitude ( Kokshoorn 2008: 99–114).