Afrotyphlops lineolatus (Jan)
Broadley, Donald G. & Wallach, Van, 2009, 2255, Zootaxa 2255, pp. 1-100 : 39-40
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publication ID |
11755334 |
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persistent identifier |
https://treatment.plazi.org/id/03E3D17B-FFD8-1258-44D4-E90D978EFAFE |
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treatment provided by |
Felipe |
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scientific name |
Afrotyphlops lineolatus (Jan) |
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( Fig. 7A)
Lineolate blind-snake
Typhlops (Ophthalmidion) lineolatus Jan, 1863: 13 . (nomen nudum)
Typhlops (Ophthalmidion) lineolatus Jan, 1864 , Icon. Gén. Ophid. I Typhl.: 24. Type locality: “ Sierra Leone,” holotype ZMUU 725, collected by A. Afzel (late 18 th –early-19 th century; lost fide Hahn, 1980: 62 ); McDiarmid et al., 1999: 107.
Typhlops lineolatus — Jan & Sordelli, 1865: 5, pl. 1, fig. 4; Günther, 1866: 24; Pitman 1938: 44, 1974: 61, pl. 1, fig. 3, pl. U, fig. 1; Roux-Estève, 1970: 2186, fig. 2, 1974: 80, fig. 13.18, 1975a: 492, fig. 30; Hahn, 1977: 37, 1980: 62; Largen, 1978: 48 (part), fig. 1; Hughes, 1983: 354; Lanza, 1983: 219, 1990: 430; Obst et al., 1984: 439, 2 figs., 1988: 774, 2 figs.; Meirte, 1992: 22, 23; Largen & Rasmussen, 1993: 323 (part); Wallach, 1993a: 217, 1994: 215, 1998a: 3, 1998b: 399; Frank & Ramus, 1995: 253; Broadley, 1998b: xii; Mattison, 1999: 147; McDiarmid et al., 1999: 107;? Kley, 2001: 1323, fig. 4; Lee & Scanlon, 2002: 400; Spawls et al., 2002: 290, fig., 2006: 90, fig.; Kley, 2003: 379, 2003: 379; Murphy, 2003: 552; Largen & Parker, 2004: 37; Brogard, 2005: 115; Lötters et al., 2007: 75, pl. 11; Conrad, 2008: 181; Cundall & Irish, 2008: 600.
Onychocepahlus lineolatus — Bocage, 1873: 250.
Typhlops (Ophthalmidion) eschrichtii lineolata — Peters, 1877: 614.
Typhlops punctatus — (not Leach) Boulenger, 1893: 42 (part); Ferreira, 1906: 167; Sternfeld, 1908a: 239 (part, Galena River at Lake Abaya); Werner, 1921: 315 (part); Schmidt, K., 1923: 45, pl. 1, fig. 1; Zavattari, 1930: 190; Curran & Kauffeld, 1937: fig. 49; Monard, 1937: 104; Fitch, 1970: 121; Porter, 1972: 406; McDiarmid et al., 1999: 107 (part).
Typhlops boulengeri Bocage, 1893 , Jorn. Sci. Math. Phys. Nat. (2) 3(10): 117. Type locality: “Quindumbo, dans l’intérieur de Benguella, Angola,” lectotype by present designation BMNH 1946.1.11.18 (formerly BMNH 1893.12.27.12) [MBL 1888 destroyed by fire 18 March 1978], collected by J. d’Anchieta; Bocage, 1895: 64, 1897: 198; Boulenger, 1896: 586, 1915a: 196; Ferreira, 1900: 50, 1906: 167; Werner, 1921: 312; Schmidt, 1923: 33; Monard, 1937: 103; Witte, 1953: 138, fig. 28, 1962: 40, 1966: 37 (part); Laurent, 1954b: 292, 1956a: 68, 366, figs. 7–8, pl. 8, fig. 3, 1956b: 282, 1960a: 18, 1960b: 439, 1964a: 89, 1965: 14; Curry-Lindahl, 1956: 41; Bourgeois, 1961: 331, 1964: 70; Thys van den Audenaerde, 1965: 369; Bourgeois, 1968: 277; Hahn, 1977: 37, 1980: 62; Derleyn et al., 1983: 771; Wallach, 1993a: 216, fig. 1B; McDiarmid et al., 1999: 93.
Typhlops punctatus lineolata — Bocage, 1895: 66.
Typhlops punctatus lineolatus — Bocage, 1895: xvii, 1896b: 112; Ferreira, 1903: 9.
Typhlops bocagei Ferreira, 1905 , Jorn. Sci. Math. Phys. Nat. Lisbon, (2) 7(26), 114. Type locality: “Cabicula, Bom Jesus (margens do Quanza)” [= Angola], syntypes (2) MBL (destroyed July 1943 during WW II), collected by Newton, 1903; Werner, 1921: 316; Monard, 1937: 103; Hahn, 1977: 37, 1980: 62; Roux-Estève, 1974: 80; McDiarmid et al., 1999: 107.
Typhlops tornieri Sternfeld, 1910b , Mitt. Zool. Mus. Berlin, 5(1): 69. Type locality: “Kilima-Ndjaro, Deutsch-Ostafrika ” [= Kilimandjaro Mountain, Tanzania], syntypes ZMB 20722–23, collected by Dr. Eggel; Sternfeld, 1910c: 11; Boulenger, 1915b: 615; Werner, 1921: 309; Schmidt, 1923: 33; Loveridge, 1924: 3, 1933: 199; Werner, 1929: 176; Roux-Estève, 1974: 80; Hahn, 1977: 37, 1980: 62; McDiarmid et al., 1999: 93; Bauer et al., 2002: 165.
Typhlops bocagii (sic) — Boulenger, 1915b: 615, 1920: 271; McDiarmid et al., 1999: 108.
Typhlops punctatus punctatus — (not Leach) Loveridge, 1929a: 16; 1933: 213; 1936b: 225; 1942: 255, 1945: 2, 1946: 6, fig. 4, 1956a: 41, 1957b: 241; Mertens, 1937: 11; Uthmöller, 1937: 100, 1941a, 26, 1941b: 227 (Kilimandjaro); Pitman, 1938: 44, pl. 1, fig. 3, pl. U, fig. 1; Laurent, 1950: 7, 1954a: 37, 1964a: 89; Witte, 1953: 139, 1966: 36; Hellmich, 1957: 70; Robertson et al., 1962: 426 (Rukwa).
Typhlops angolensis boulengeri — (not Bocage) Laurent, 1964b: 392.
Typhlops boulengeri boulengeri — (not Bocage) Laurent, 1964b: 414; Guibé in Pitman, 1974: 59; McDiarmid et al., 1999: 93.
Typhlops lineolatus lineolatus — Roux-Estève, 1974: 80, 90, 124, figs. 13.18 & 54; Largen, 1978: 51; Poynton & Broadley, 1978: 930; Spawls, 1978: 2; Hahn, 1980: 62; Rodgers & Homewood, 1982: 238; Welch, 1982: 129, 1994: 71; Pakenham, 1983: 32; Broadley, 1991: 525; Broadley & Howell, 1991: 21; Rasmussen, 1991: 181, fig. 2; Simbotwe & Mubamba, 1993: 76; Broadley, 1998a: 38; McDiarmid et al., 1999: 108.
Typhlops boulengerii — Roux-Estève, 1974: 80; Largen, 1978: 48.
Typhlops eschrichtii —(not Schlegel) Largen, 1978: 48.
Typhlops angolensis blanfordi — Hahn, 1980: 62; McDiarmid et al., 1999: 108.
Typhlops lineolatus tanganicanus — Hahn, 1980: 62 (part).
Typhlops schmidti schmidti — Hahn, 1980: 62.
Typhlops lincelatus (sic)— Chifundera, 1990: 141.
Rhinotyphlops lineolatus — Broadley & Wallach, 2000: 167; Broadley & Cotterill, 2004: 44.
Description. Snout rounded, prominent. Rostral broad, three-quarters the width of the head, truncated posteriorly; frontal subtrapezoidal; supraocular oblique, its lateral apex between nasal and preocular (rarely between preocular and ocular); eye usually behind the posterior border of the preocular; nasal suture arising from the first labial; SIP usually T-X (P, P, O, O), sometimes II-O, very rarely I-P; MSR 26 to 30, reduction A/B +2 to 2, B/C 2 to 6; MD 323–505 (mean 379.1, n = 169); vertebrae 185–225 (mean 201.2, n = 82); MD/V ratio 1.67–2.05 (mean 1.87, n = 82); L/D ratio 21–47 (mean 31.8, n = 114). Blackish, each scale with yellow spots at base and apex, pigmentation gradually reduced laterally and sometimes with the midventral zone unpigmented or with a series of irregular pale patches. MCZ 30037 from Ujiji is black above and blotched below. Largest specimen (IRSNB 3206A—Lubumbashi, Democratic Republic of the Congo) has a total length of 640 mm.
Distribution. Savannas of West Africa, east to Ethiopia, south to Angola, the Katanga Province of the Democratic Republic of the Congo, and northern Tanzania, 0–2600 m ( Figs. 11, 13 & 14).
Localities. SUDAN. Kagelu MCZ 45261 ; Kajiko North ZFMK 26061 View Materials ; Katiri MCZ 53321–22 View Materials ; Suni, Darfur Mts ZFMK 39898–99 View Materials ; Talanga Forest, Imatong Mts ZFMK 26062 View Materials , 29558 View Materials , 29687–93 View Materials , 29708–10 View Materials , 297220– 26 View Materials , 29748–52 View Materials ; Yambio BMNH 1951.1.7.86–7 . ETHIOPIA. Galena River at Lake Abaya ZMB 27453 . DEM- OCRATIC REPUBLIC OF THE CONGO. (Orientale Province) Aru Kilabi IRSNB 3228 View Materials ; MRAC 1232 View Materials ; Avakubi BMNH 1919.8.16.73–74; MRAC 1614 View Materials (5) ; Bagbele IRSNB 3107–08 View Materials (2) ; Ekibondo ANSP 20500 View Materials ; Faradje AMNH 11609–11 View Materials , 11618–20 View Materials ; IRSNB 3969 View Materials ; Gangala na Bodio IRSNB 3103 View Materials , 3110 View Materials ; Garamba AMNH 11612–17 View Materials ; IRSNB 3102 View Materials , 3104–06 View Materials , 3109 View Materials ; Mahagi MRAC 10161 View Materials . (Kivu Province) Kagondo MRAC 20342– 43 View Materials ; Kiharo MRAC 17613–15 View Materials ; Kitutu MRAC 21629 View Materials ; Lubirizi MCZ 57457, MRAC 20655–95 View Materials ; Mahagi-Port, Lac Albert MRAC 20627 View Materials ; Makobola MRAC 21243 View Materials ; Mugera MRAC 17609–12 View Materials ; Rugari MRAC 10919 View Materials ; Runingo MRAC 20257–59 View Materials ; Uvira MNHN 1966-1239 View Materials - 43 View Materials , MRAC 17616–33 View Materials , 20389 View Materials , 20655–93 View Materials , 20793–796 View Materials , 21283–300 View Materials , 21342 View Materials , 21537–55 View Materials . (Katanga Province) Bukena IRSNB 4001 View Materials A; Kalémie IRSNB 3990 View Materials ; MRAC 610 View Materials ; Kapiri IRSNB 3975 View Materials ; Likasi IRSNB 3977 View Materials ; Lubumbashi IRSNB 3206 View Materials (2) ; Musosa IRSNB 3967–68 View Materials ; Panda IRSNB 3976 View Materials , 3978–79 View Materials (2) ; Upemba National Park IRSNB 3992–96 View Materials , 4000 View Materials . (Bas-Zaire Province) Kisantu Botanical Gardens BMNH 1980.29 – 30 . UGANDA. Ajai Game Reserve MUZM —; Budongo Forest BMNH 1935.10 . 11.2, 1936.7.3.3–5, LACM 35282–83 View Materials ; Bugiri ( Bourgeois, 1961); Bulaba BMNH 1954.1.11.87; Chagwe ZMB 58377 ; Jinja BMNH 1912.10.12.1; MCZ 30042 ; Kampala MUZM —; Kitwanga BMNH 1912.10.14.3; Kome Island BMNH 1936.7.3.6; Lukambi BMNH 1934.12.15.519; Mabira Forest BMNH 1908.10.20.9–10; CAS 111849 View Materials , MCZ 30040–41 View Materials , 48059–60 View Materials , NMZB 127–28 View Materials ; Masindi NMZB 125 View Materials ; Mityama NMK 352–53 View Materials ; Mubango BMNH 1934.12.15.20; Murchison Falls BMNH 1908.2.4.2; Mt. Elgon BMNH 1901.10.14.1–2; Nyakabanda USNM 120797 View Materials . KENYA. Athi River NMK 2692 View Materials ; Chapelunge MMK 880 View Materials ; Chemelil CAS 148005 View Materials ; Embakasi NMK 2782 View Materials ; Juja Farm USNM 40898 View Materials ; Kakamega MCZ 96853 ; NMK 2306 View Materials (2) ; Kavirondo Gulf ZMB 11922 ; Kiambu NMK 149 View Materials ; Kiboko Range NMK 2948 View Materials ; Kisumu NMK 3356 View Materials , 3368 View Materials ; NMZB-UM 8958–9 ; Kitale NMK 1503 View Materials , 2975 View Materials ; Lolgorien NMK 3106 View Materials ; Makueni NMK 1584 View Materials ; Masai Mara Game Reserve NMZB 7915 View Materials ; Meru TMP 26561 ; Nairobi NMK 150–2 View Materials , 156–7 View Materials ; Nyambeni Hills NMK 2008 View Materials ; Yala River NMK 3356 View Materials , 3368 View Materials . TANZANIA. Amani NMZB 16145 View Materials ; Handeni-Yamota MCZ 54811 ; Ikiju- Musoma MCZ 54808 ; Imanina Hills BMNH 1959.1.4.65; Kasulu MCZ 54651 ; Kibondo NMZB 6957 View Materials , 7434 View Materials ; Kibonoto MCZ 38684 ; Kigoma BMNH 1959.1.4.58; MCZ 54810 ; Kilimandjaro ZMU 20722–3 (syntypes of Typhlops tornieri Sternfeld ); Kilombero Valley KMH 17651, 17671; NMZB 17341 View Materials ; Lake Manyara National Park NMZB 8329 View Materials ; Magrotto Hill MCZ 48061, 48063–5 View Materials ; MRAC 21947 View Materials ; NMZB 14080–81 View Materials ; Majita — Musoma MCZ 54809 ; Muse, N. Rukwa MCZ 54586 ; Mwanza BMNH 1976.2294 , MCZ 30038 ; Ngorongoro ( Bourgeois, 1961); Rukwa Valley IRSNB 3229 View Materials , 3232 View Materials ; Sekenke — Somanza MCZ 49507 ; MRAC 21948 View Materials ; Tanga MNHN 1897-233 View Materials ; Ujiji MCZ 30037 ; Ukerewe Island MCZ 30039 ; Uluguru North NMZB 17549 View Materials ; Zaraninge Forest Reserve NMZB 14523 View Materials . ANGOLA. Cazengo ( Ferreira, 1903); Quindumbo BMNH 1946.1 . 11.18 (lectotype of Typhlops boulengeri Bocage ); Rio Luinha ( Ferreira, 1906) .
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Afrotyphlops lineolatus (Jan)
| Broadley, Donald G. & Wallach, Van 2009 |
Rhinotyphlops lineolatus
| Broadley, D. G. & Cotterill, F. P. D. 2004: 44 |
| Broadley, D. G. & Wallach, V. 2000: 167 |
Typhlops lincelatus
| Chifundera, K. 1990: 141 |
Typhlops (Ophthalmidion) lineolatus
| McDiarmid, R. W. & Campbell, J. A. & Toure, T. A. 1999: 107 |
| Hahn, D. E. 1980: 62 |
Typhlops angolensis blanfordi
| McDiarmid, R. W. & Campbell, J. A. & Toure, T. A. 1999: 108 |
| Hahn, D. E. 1980: 62 |
Typhlops lineolatus tanganicanus
| Hahn, D. E. 1980: 62 |
Typhlops schmidti schmidti
| Hahn, D. E. 1980: 62 |
Typhlops eschrichtii
| Largen, M. J. 1978: 48 |
Typhlops lineolatus lineolatus
| McDiarmid, R. W. & Campbell, J. A. & Toure, T. A. 1999: 108 |
| Broadley, D. G. 1998: 38 |
| Welch, K. R. G. 1994: 71 |
| Simbotwe, M. P. M. & Mubamba, R. 1993: 76 |
| Broadley, D. G. & Howell, K. M. 1991: 21 |
| Rasmussen, J. B. 1991: 181 |
| Pakenham, R. H. W. 1983: 32 |
| Rodgers, W. A. & Homewood, K. M. 1982: 238 |
| Welch, K. R. G. 1982: 129 |
| Hahn, D. E. 1980: 62 |
| Largen, M. J. 1978: 51 |
| Poynton, J. C. & Broadley, D. G. 1978: 930 |
| Spawls, S. 1978: 2 |
| Roux-Esteve, R. 1974: 80 |
Typhlops boulengerii
| Largen, M. J. 1978: 48 |
| Roux-Esteve, R. 1974: 80 |
Typhlops angolensis boulengeri
| Laurent, R. F. 1964: 392 |
Typhlops boulengeri boulengeri
| McDiarmid, R. W. & Campbell, J. A. & Toure, T. A. 1999: 93 |
| Pitman, C. R. S. 1974: 59 |
| Laurent, R. F. 1964: 414 |
Typhlops punctatus punctatus
| Witte, G. - F. de 1966: 36 |
| Laurent, R. F. 1964: 89 |
| Robertson, I. A. D. & Chapman, B. M. & Chapman, R. F. 1962: 426 |
| Hellmich, W. 1957: 70 |
| Laurent, R. F. 1954: 37 |
| Witte, G. - F. de 1953: 139 |
| Laurent, R. F. 1950: 7 |
| Loveridge, A. 1945: 2 |
| Loveridge, A. 1942: 255 |
| Pitman, C. R. S. 1938: 44 |
| Mertens, R. 1937: 11 |
| Uthmoller, W. 1937: 100 |
| Loveridge, A. 1936: 225 |
| Loveridge, A. 1933: 213 |
| Loveridge, A. 1929: 16 |
Typhlops bocagei
| McDiarmid, R. W. & Campbell, J. A. & Toure, T. A. 1999: 107 |
| Hahn, D. E. 1980: 62 |
| Hahn, D. E. 1977: 37 |
| Roux-Esteve, R. 1974: 80 |
| Monard, A. 1937: 103 |
| Werner, F. 1921: 316 |
Typhlops bocagii
| McDiarmid, R. W. & Campbell, J. A. & Toure, T. A. 1999: 108 |
| Boulenger, G. A. 1920: 271 |
| Boulenger, G. A. 1915: 615 |
Typhlops tornieri
| McDiarmid, R. W. & Campbell, J. A. & Toure, T. A. 1999: 93 |
| Hahn, D. E. 1980: 62 |
| Hahn, D. E. 1977: 37 |
| Roux-Esteve, R. 1974: 80 |
| Loveridge, A. 1933: 199 |
| Werner, F. 1929: 176 |
| Loveridge, A. 1924: 3 |
| Schmidt, K. P. 1923: 33 |
| Werner, F. 1921: 309 |
| Boulenger, G. A. 1915: 615 |
| Sternfeld, R. 1910: 11 |
Typhlops punctatus lineolatus
| Ferreira, J. de & Bethencourt 1903: 9 |
Typhlops boulengeri
| McDiarmid, R. W. & Campbell, J. A. & Toure, T. A. 1999: 93 |
| Wallach, V. 1993: 216 |
| Derleyn, P. & Taverne, L. & Ndabakubije, V. & Madodo, G. & Karonkano, J. 1983: 771 |
| Hahn, D. E. 1980: 62 |
| Hahn, D. E. 1977: 37 |
| Bourgeois, M. 1968: 277 |
| Thys van den Audenaerde, D. F. E. 1965: 369 |
| Bourgeois, M. 1964: 70 |
| Bourgeois, M. 1961: 331 |
| Laurent, R. F. 1956: 68 |
| Curry-Lindahl, K. 1956: 41 |
| Laurent, R. F. 1954: 292 |
| Witte, G. - F. de 1953: 138 |
| Monard, A. 1937: 103 |
| Schmidt, K. P. 1923: 33 |
| Werner, F. 1921: 312 |
| Ferreira, J. de & Bethencourt 1906: 167 |
| Ferreira, J. de & Bethencourt 1900: 50 |
| Bocage, J. V. 1897: 198 |
| Bocage, J. V. 1895: 64 |
Typhlops punctatus lineolata
| Bocage, J. V. 1895: 66 |
Typhlops punctatus
| McDiarmid, R. W. & Campbell, J. A. & Toure, T. A. 1999: 107 |
| Porter, K. R. 1972: 406 |
| Fitch, H. S. 1970: 121 |
| Monard, A. 1937: 104 |
| Zavattari, E. 1930: 190 |
| Werner, F. 1921: 315 |
| Sternfeld, R. 1908: 239 |
| Ferreira, J. de & Bethencourt 1906: 167 |
| Boulenger, G. A. 1893: 42 |
Typhlops (Ophthalmidion) eschrichtii lineolata
| Peters, W. C. H. 1877: 614 |
Onychocepahlus lineolatus
| Bocage, J. V. 1873: 250 |
Typhlops lineolatus
| Conrad, J. L. 2008: 181 |
| Cundall, D. & Irish, F. 2008: 600 |
| Lotters, S. & Wagner, P. & Bwong, B. A. & Schick, S. & Malonza, P. K. & Muchai, V. & Wasonga, D. V. & Veith, M. 2007: 75 |
| Brogard, J. 2005: 115 |
| Largen, M. J. & Parker, A. R. 2004: 37 |
| Kley, N. J. 2003: 379 |
| Kley, N. J. 2003: 379 |
| Murphy, J. B. 2003: 552 |
| Lee, M. S. Y. & Scanlon, J. D. 2002: 400 |
| Spawls, S. & Howell, K. M. & Drewes, R. C. & Ashe, J. 2002: 290 |
| Kley, N. J. 2001: 1323 |
| Mattison, C. 1999: 147 |
| McDiarmid, R. W. & Campbell, J. A. & Toure, T. A. 1999: 107 |
| Wallach, V. 1998: 3 |
| Wallach, V. 1998: 399 |
| Frank, N. & Ramus, E. 1995: 253 |
| Wallach, V. 1994: 215 |
| Largen, M. J. & Rasmussen, J. B. 1993: 323 |
| Wallach, V. 1993: 217 |
| Meirte, D. 1992: 22 |
| Lanza, B. 1990: 430 |
| Hughes, B. 1983: 354 |
| Lanza, B. 1983: 219 |
| Hahn, D. E. 1980: 62 |
| Largen, M. J. 1978: 48 |
| Hahn, D. E. 1977: 37 |
| Pitman, C. R. S. 1974: 61 |
| Roux-Esteve, R. 1970: 2186 |
| Pitman, C. R. S. 1938: 44 |
| Gunther, A. C. L. G. 1866: 24 |
| Jan, G. & Sordelli, F. 1865: 5 |
Typhlops (Ophthalmidion) lineolatus
| Jan, G. 1863: 13 |