Agallamyia Norrbom, 2010

Norrbom, Allen L., Sutton, Bruce D., Steck, Gary J. & Monzón, José, 2010, New genera, species and host plant records of Nearctic and Neotropical Tephritidae (Diptera) 2398, Zootaxa 2398, pp. 1-65 : 12-14

Agallamyia Norrbom , new genus

Type species: A. pendula Norrbom , new species.

Diagnosis. Agallamyia resembles Pyrgotoides Curran in general habitus and in having a setulose lunule, but differs as follows: Wing ( Fig. 33) with only narrow irregular hyaline streaks in addition to hyaline triangle on anterior margin distal to apex of vein R 1, without large hyaline areas on posterior margin as in Pyrgotoides ; male fore femur ( Fig. 14) very stout, with row of ventral setae stout and spinelike (the male fore femur is sometimes stout in Pyrgotoides , but the ventral setae are more slender and not spinelike); tibiae not exceptionally stout (often very stout in Pyrgotoides ); and eversible membrane ( Fig. 15) similar dorsally and ventrally, both sides with broad medial area of spinules with sharply delimited transition from larger to minute spinules (ventral stripes of spinules extended from the taenia separate, not fused to form single narrow medial stripe as in Pyrgotoides ). Based on the difference in the eversible membrane, the similarities in external characters between Agallamyia and Pyrgotoides are probably due to convergence. The fusion of the ventral stripes of spinules on the eversible membrane as in Pyrgotoides was considered a synapomorphy of the Dithrycini by Korneyev (1999), although his classification of this group was not supported by Han et al. (2006). Unless the condition in Agallamyia is secondarily derived, Agallamyia appears to belong to the Cecidocharini rather than the Dithrycini . Agallamyia is probably most closely related to Ostracocoelia Giglio-Tos , which has a similar eversible membrane, short thoracic setae, stout, ventrally spinose male fore femur, and sometimes setulose lunule (in Ostracocoelia the lunule is never as densely setulose as in Agallamyia , but often has one to several fine setulae). All of these characters are apomorphic and probably are synapomorphies for these two genera. Agallamyia differs from Ostracocoelia in wing pattern (not banded), thoracic microtrichia (without large shiny nonmicrotrichose areas on anepisternum and scutellum as in Ostracocoelia ), chaetotaxy (e.g., presutural supra-alar seta absent, scutellum setulose), and eye size (genal height more than half long diameter of eye vs. less than half (0.29–0.35) in Ostracocoelia ).

Description. Relatively large, brown. Setae short.

Head ( Figs. 11–12): Frons broad, nonsetulose anteromedially. Setae relatively short; 2 orbital setae; medial and lateral vertical setae subequal; postocular setae acuminate, subequal, yellow. Face with broad carina. Lunule large and setulose. Eye relatively small.

Thorax ( Fig. 13): With following setae short (length less than distance between acrostichal setae): postpronotal, 2 notopleural, usually 1 postsutural supra-alar (absent in 2 specimens and on 1 side in 2 others; when present relatively posterior in position, at 1/2–3/4 distance from transverse suture to posterior margin of scutum), 1 intra-alar, 1 postalar, 0–1 dorsocentral (aligned with postsutural supra-alar seta to midway between it and postalar seta), 1 acrostichal, and 2 scutellar; anepisternal and anepimeral setae sometimes poorly or not differentiated from setulae; presutural supra-alar seta and katepisternal seta absent. Mesonotum without shiny, nonmicrotrichose area. Scutellum short and convex.

Legs: Stout. Male fore femur with row of ventral setae stout and spinelike. Hind femur with preapical setae undifferentiated.

Wing ( Fig. 33): Elongate and relatively narrow. Pterostigma short and narrow. Subcosta with apical part weak, turned anterodistally at 45° angle. Costal setulae at subcostal break undifferentiated, no larger than other costal setulae. Vein R 1 dorsally without gap in setulae near bend in vein Sc. Vein R 4+5 dorsally intermittently setulose. Lobe of cell bcu moderately long. Crossvein r-m distinctly distal to midlength of cell dm. Pattern predominantly brown, with only narrow irregular hyaline to yellowish streaks in addition to hyaline triangle on anterior margin distal to apex of vein R 1.

Abdomen: Tergites without shiny nonmicrotrichose areas.

Male terminalia: Surstyli short. Glans with relatively stout sclerotized part including laterally curved acrophallus.

Female terminalia: Eversible membrane ( Fig. 15) similar dorsally and ventrally, with pair of short taenia extended to single broad oval medial area of spinules with sharply delimited transition from larger spinules anteriorly and narrowly laterally to minute spinules medially and posteriorly. Aculeus with slender, simple tip.

Etymology. This name is derived from the Spanish “agalla” (gall) and Greek “myia” (fly). It is considered feminine in gender.