Bothynus entellus

Duarte, Paulo R. M. & Grossi, Paschoal C., 2020, Bothynus entellus (LePeletier & Serville) (Coleoptera: Scarabaeidae: Dynastinae) species group: taxonomic revision and description of two new species, Zootaxa 4750 (1), pp. 101-121 : 120

publication ID	https://doi.org/ 10.11646/zootaxa.4750.1.5
publication LSID	lsid:zoobank.org:pub:1A3B7CFF-292B-46AB-82E8-C78BA34618EC
DOI	https://doi.org/10.5281/zenodo.3706219
persistent identifier	https://treatment.plazi.org/id/03E31B7A-5911-FFB7-FF6C-FF4FBB8AF89D
treatment provided by	Plazi
scientific name	Bothynus entellus
status

Treatment

Key to males of Bothynus entellus species group

Males have pronotal horns, pronounced pronotal excavations, and enlarged anterior claws.

1. Pronotum with 2–3 horns ( Figs. 1A View FIGURE 1 , 3A View FIGURE 3 ), or only with a small ( Fig. 5A View FIGURE 5 ) to moderate ( Fig. 2A View FIGURE 2 ) apical tubercle; if moderate, apical tubercle with rounded apex; mesofemora with setae confine on disc ( Fig. 5F View FIGURE 5 ); tergite VII with 2 bands of weakly marked carinae ( Fig. 7A View FIGURE 7 ); parameres without laterovental contraction at apical half ( Fig. 9A View FIGURE 9 )............................... 2

- Pronotal tubercle strong, horn shaped, acute on apex ( Fig. 4A View FIGURE 4 ); mesofemora completely setose ( Fig. 4B View FIGURE 4 ); tergite VII with 1 band of well-marked carinae ( Fig. 7C View FIGURE 7 ); parameres with lateroventral contraction at apical half in lateral view ( Fig.9D View FIGURE 9 )................................................................................ Bothynus lancifer Dechambre, 1981

2. Pronotum strongly convex posteriorly in lateral view ( Fig. 2B View FIGURE 2 ); pronotum only with an apical tubercle moderate in size; protarsomere IV with elongate ventral apex nearly reaching the apex of the ventro-basal tooth of tarsomere V ( Fig. 2E View FIGURE 2 )......... 3

- Pronotum moderately convex ( Fig. 5B View FIGURE 5 ), or with a horn ( Figs. 1A View FIGURE 1 , 3A View FIGURE 3 ) or tubercle ( Figs. 1C View FIGURE 1 , 3C View FIGURE 3 ) on each side); protarsomere IV with short ventral apex, not reaching the base of the ventro-basal tooth of tarsomere V ( Fig. 5E View FIGURE 5 ).................... 4

3. Metatrochanter with rounded apex ( Fig. 2F View FIGURE 2 ); parameres with subparallel outer margins ( Fig. 8B View FIGURE 8 )............................................................................................ Bothynus stenelus ( Burmeister, 1847)

- Metatrochanter with acute apex ( Fig. 6B View FIGURE 6 ); parameres with rounded lateral flaps ( Fig. 8F View FIGURE 8 )........................................................................................ Bothynus condacki Duarte & Grossi , new species

4. Pronotum with moderate ( Fig. 1A View FIGURE 1 ) or shallow ( Fig. 5A View FIGURE 5 ) concavity; hypomeron not emarginate in ventral view ( Fig. 5D View FIGURE 5 )... 5

- Pronotal concavity extremely deep ( Fig. 3A View FIGURE 3 ); hypomeron strongly emarginate in ventral view ( Fig. 3E View FIGURE 3 )............................................................................................ Bothynus tricornis Arrow, 1937

5. Stridulatory apparatus with well-marked carinae near to basal margin ( Fig. 7B View FIGURE 7 ); apical half of parameres with subparallel lateral flaps, narrower than basal half ( Fig. 8E View FIGURE 8 ).............................. Bothynus araya Duarte & Grossi , new species

- Stridulatory apparatus with weakly marked carinae near to basal margin ( Fig. 7A View FIGURE 7 ); apical half of parameres with rounded lateral flaps, broader than basal half ( Fig. 8A View FIGURE 8 ).............................. Bothynus entellus ( LePeletier & Serville, 1828)