Binaluana, Soulier-Perkins, Adeline & Stroiński, Adam, 2015

Soulier-Perkins, Adeline & Stroiński, Adam, 2015, A new lophopid genus as another piece in the biogeographical history puzzle of the family in the Sunda Shelf (Hemiptera: Fulgoromorpha: Lophopidae), Zootaxa 4006 (3), pp. 586-600 : 587-593

publication ID

https://doi.org/ 10.11646/zootaxa.4006.3.10

publication LSID

lsid:zoobank.org:pub:5169CFEE-071C-40F1-9A16-4E898975EA33

DOI

https://doi.org/10.5281/zenodo.6122419

persistent identifier

https://treatment.plazi.org/id/03E1BE04-F038-FFFD-FF7D-B59FFD63F82C

treatment provided by

Plazi

scientific name

Binaluana
status

gen. nov.

Binaluana gen. nov.

( Figs 1–60 View FIGURES 1 – 5 View FIGURES 6 – 11 View FIGURES 12 – 17 View FIGURES 18 – 23 View FIGURES 24 – 29 View FIGURES 30 – 35 View FIGURES 36 – 41 View FIGURES 42 – 47 View FIGURES 48 – 52 View FIGURES 53 – 60 )

Type species: Binaluana emarginata sp. nov., here designated.

Etymology. Melichar intended to name this genus after the location where it was found, Binaluan. We kept his name Binaluana . Gender: feminine.

Diagnosis. Binaluana gen. nov. similar to Zeleja and Bisma , but Bisma presents some carinae on the frons, when frons in Binaluana and Zeleja presents none. Binaluana and Bisma have a shorter prothorax than mesothorax, with a clear produced disc, when Zeleja has a pro- and mesothorax equal in length.

Description. Body elongate and narrow ( Figs. 1–2 View FIGURES 1 – 5 ).

HEAD. Head with compound eyes narrower than prothorax ( Figs 2, 5 View FIGURES 1 – 5 & 12 View FIGURES 12 – 17 ). Vertex longer than wide, flat, without median carina, lateral margins carinate, parallel and elevated, anterior margin obsolete ( Figs 4 View FIGURES 1 – 5 , 15–17 View FIGURES 12 – 17 ). Frons generally flat with the exception of the protuberance just bellow the upper margin, no carinae, lateral margins carinate and continuous from the vertex to the frontoclypeal margin. Frons bottle-shape, with widest part along the frontoclypeal suture. In lateral view, top of the head higher than the surface of pro- and mesothorax, compound eyes rounded, flattened postero-ventrally, with margin cut out ventrally; ocelli present; foramen of antenna not touching the compound eyes; ocellar and genal carina absent ( Figs 1, 3 View FIGURES 1 – 5 , 13 View FIGURES 12 – 17 , 18–22 View FIGURES 18 – 23 ). Carinated line starting at the edge of frontoclypeal suture and lateral frontal margin, running toward the ocellar and disappearing ( Figs 1–2 View FIGURES 1 – 5 , 18– 22 View FIGURES 18 – 23 ). Pedicel of antenna twice longer than wide ( Figs 4–5 View FIGURES 1 – 5 , 24–27 View FIGURES 24 – 29 ). Sensory plate organs multi-petals shaped, surrounded by strong denticules, present on whole pedicel, arranged in lines ( Figs 24–29 View FIGURES 24 – 29 ); trichoid sensilla present (type 1 and 2). Clypeus strongly carinate laterally without median carina ( Figs 4 View FIGURES 1 – 5 , 6 View FIGURES 6 – 11 & 15 View FIGURES 12 – 17 ). Rostrum reaching metacoxa, apical segment distinctly shorter than subapical one ( Figs 6 View FIGURES 6 – 11 & 41 View FIGURES 36 – 41 ).

THORAX. Pronotum anteriorly rounded, median disc tricarinate, carinae connected anteriorly and reaching posterior margin ( Figs 2, 5 View FIGURES 1 – 5 , 12–13 View FIGURES 12 – 17 ). Median disc elevated and flattened, slightly protruding anteriorly. Strong carina present on lateral part of pronotum ( Figs 18–19, 21–22 View FIGURES 18 – 23 ). Posterior margin widely rounded. Mesothorax with a single median carina starting on the anterior margin but not reaching the scutelum. Pro- and mesothoracic legs flattened, but not foliaceous, with femur and tibia of the same length, prothoracic leg more flattened than mesothoracic one ( Figs 6 View FIGURES 6 – 11 , 36 View FIGURES 36 – 41 ). Metathoracic leg with tibia longer than femur. Tibia with 2 lateral spines on the lower half, row of large apical teeth present, with median diastemma. First tarsal segment bearing apical spines organised in a triangular area, second tarsal segment without any spine reduced into a small rounded lobe ( Figs 7 View FIGURES 6 – 11 , 37–41 View FIGURES 36 – 41 ).

TEGMINA ( Figs 1–2 View FIGURES 1 – 5 , 30–35 View FIGURES 30 – 35 ). Tegmen elongate, flattened, membranous, completely covered with setae, on and between veins. Costal margin almost parallel to postclaval margin. Costal area narrow at the base and getting wider toward the apex, reaching the level of claval apex, transversal veinlets present all along it. Postcostal cell with veinlets. Basal cell much narrower than long. ScP+R+MA leaving basal cell by a long common stem. Forking of the veins, leaving the basal cell, is done in the following order: ScP+R+MA, CuA and MP. ScP+RA forking almost at same level than secondary forking of RP and both a little before the end of costal area. MP forking before ScP+RA forking. MP1+2 forking before MP3+4 and not forking any further. CuA forking a little later than fusion of Pcu+A1, all CuA terminals reaching postclaval margin. Transverse veinlets present in radial and median cells, but weakly visible. Few transverse veinlets present in apex of cubital cell. Transverse veinlets between CuP and Pcu and Pcu+A1.

MALE. Unknown.

FEMALE TERMINALIA. Seventh abdominal sternite bearing a protuberance ventrally ( Figs 7, 10 View FIGURES 6 – 11 , 44–45 View FIGURES 42 – 47 , 48 View FIGURES 48 – 52 –50, 53–55 View FIGURES 53 – 60 ), anterior margin convex anteriorly and posterior margin with lateral corners protruding. Medio-ventral extension at the apex of the anal tube absent. Anal tube bearing two large lobes apically, oriented postero-ventrally and covering the gonoplacs ( Figs 9–11 View FIGURES 6 – 11 , 42–47 View FIGURES 42 – 47 , 56 View FIGURES 53 – 60 ). Small longitudinal carina along the ventral part of anal tube. Dense and long setae covering the anal tube.

Gonoplacs, unilobate elongate-rounded and bearing dense setae internally and externally ( Figs 42–45 View FIGURES 42 – 47 , 52 View FIGURES 48 – 52 , 57 View FIGURES 53 – 60 ). Gonocoxae IX present, gonapophysis IX clearly visible when for the other Lophopid genera they are strongly reduced and hardly observable sometime ( Fig. 57 View FIGURES 53 – 60 ). Bursa copulatrix with a single pouch with visible cells ( Fig. 58 View FIGURES 53 – 60 ).

Spermatheca well developed, ductus receptaculi distinctly longer than diverticulum ductus, ribbed. Diverticulum ductus smooth, bulbous ( Figs 59–60 View FIGURES 53 – 60 ).

As for all Lophopidae , gonospiculum absent.

Distribution. Palawan (The Philippines).

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Lophopidae

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