Evansiclavelia poecilopteryx Pitts, Rodriguez & Shimizu, 2021

Shimizu, Akira, Pitts, James P, Rodriguez, Juanita, Wahis, Raymond & Yoshimura, Jin, 2021, Systematics and convergent evolution in three Australian genera of Pepsinae spider wasps (Hymenoptera: Pompilidae), Austral Entomology 60 (2), pp. 301-316 : 11-13

publication ID

https://doi.org/ 10.1111/aen.12530



persistent identifier


treatment provided by


scientific name

Evansiclavelia poecilopteryx Pitts, Rodriguez & Shimizu

sp. nov.

Evansiclavelia poecilopteryx Pitts, Rodriguez & Shimizu , sp. nov.

( Fig. 4 View Fig ) http://zoobank.org/urn:lsid:zoobank.org:act:F1D50B45-B938- 4376-B037-E4F7A0336C74

Material examined

Holotype ♀ ‘ QLD Cape York Heathlands RP 11.616° 142.80930° GE 29 Nov 2017 Creekbed Cpt Billy’ s Landing Rd J A & J G Lumbers’ ‘ PO1486 ’ ‘Maurillus’ ‘Molecular Voucher Pitts 2019 UCE PO1486’ ( ANIC).



Body and legs black; head, pronotum and mesonotum wine red ( Fig. 4c,d,h,i View Fig ). Wings transparent with dark brown fascia occupying basal half of marginal cell, SMC1–3, discal cells 1 and 2, subdiscal cell 1, basal two-thirds of subdiscal cell 2, and clavus apically ( Fig. 4j View Fig ).



Based on holotype.

Length: body 8.2 mm; fore wing 7.4 mm. Following wine red: head, scape, mouthparts, pronotum, mesonotum, propleuron, mesopleuron anterodorsally and posteroventrally, fore coxa basally and apically, mid and hind coxae apically, fore trochanter mostly, mid and hind trochanters apically, fore and mid femora apically, fore tibia and tarsomeres 1–4 except dorsally, fore tarsomere 5, and tarsal claws apically. Following light yellowish brown: tegula, wing bases, posterolateral margin of metanotum, fore and mid tibial spurs, metasomal terga laterally and posteriorly (laterotergite reddish yellow), and S1–5 posteriorly.

Upper frons, vertex, pronotum and mesonotum with short coppery pubescence. Lower frons, clypeus, gena, pronotum ventrolaterally, propleuron, mesopleuron, metapleuron, mesosternum, metapostnotum laterally, propodeum laterally and posteriorly, legs, and metasomal terga with silvery-white pubescence long and dense on propleuron, mesosternum, propodeum posteriorly, coxae, and T1–5 posterolaterally. Propleuron, propodeum posterolaterally, fore coxa, T1 basally, T6 and S6 with long, somewhat yellowish pale setae.

Frons, vertex and clypeus polished with small punctures irregular in size and spacing. Pronotum and mesonotum minutely tessellate and matte; mesoscutum medially and scutellum with small, dense punctures. Metasomal terga polished ( Fig. 4i View Fig ). S1–5 strongly smooth and polished with small punctures sparsely. S6 irregularly and strongly punctate.

Head 1.2× as broad as high ( Fig. 4a View Fig ). Vertex raised between and behind posterior ocelli; posterior margin broadly and strongly emarginate in dorsal view ( Fig. 4b View Fig ). Frons flattened at lower half; frontal line almost vestigial. Antennocular line slightly convex. Inner orbits barely emarginate above middle, divergent below in whole ( Fig. 4a View Fig ). UID:MID: LID = 7.9:10:10. MID 0.48× TFD. Eye width 1.1× half of MID. POL:OOL = 1:0.64. Ocelli forming right-angled triangle ( Fig. 4b View Fig ); OOcD/POD = 2.1. Clypeus width 1.6× length. Malar space longer than pedicel width. Labrum not bent backward from clypeus. Gena width ~0.3× eye width ( Fig. 4c View Fig ). Mandible with anterior face flattened; inner margin sharply edged with small tooth near apical point; outer margin with broad lamina delimited by fimbriate groove from major part. Antenna rather long and thin. Scape:pedicel:fl:fl2 = 0.81:0.25:1:0.81. Fl1 4.6× as long as wide, 0.92× UID.

Pronotum short with posterior margin arcuate ( Fig. 4b View Fig ). Mesoscutum convex; posterolateral margin narrowly and sharply reflexed. Discs of scutellum slightly convex withnarrow, blunt median carina ( Fig. 4e View Fig ). Metapostnotum very short, ~0.1× as long as metanotum at midline, deeply depressed between metanotum and propodeum. Propodeum in dorsal view 1.1× as wide as long; dorsum strongly rugose and slightly bumped posteromedially ( Fig. 4e View Fig ), forming angle of ~120° with declivity but not clearly divided from latter; median groove indistinct; sides almost parallel-sided; declivity longer than dorsum with fine transverse rugae.

Hind tibial spur 0.48× as long as hind tarsomere 1.

Fore and hind wings as shown in Fig. 4j View Fig . Marginal cell removed from wing tip by 0.6× its own length. SMC2: SMC3 = 1:1.2 on vein M, 1:0.53 on vein Rs. SMC2 narrowed on vein Rs by 0.9× its length on vein M, receiving crossvein 1m-cu at basal 0.45. SMC3 narrowed on vein Rs by 0.4× its length on vein M, receiving crossvein 2m-cu at basal 0.79. Crossveins 2rs-m nearly straight. Crossvein 3rs-m barely curved. Crossvein cu-a originating slightly distal to point of separation of vein M + CuA, weakly curved. Hind wing crossvein rs-m straight, nearly perpendicular to vein M. Crossvein cu-a originating slightly proximal to separation of vein M + CuA.

Metasoma spindle-shaped ( Fig. 4i View Fig ). T1 very shortly parallel-sided basally, with anterior face not flattened or vertical.




Cape York, Queensland, Australia.


The species name is derived from the transparent fore wing with subapical broad dark brown fascia.


Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection