Telmatoscopus advena ( Eaton, 1893 )

Kvifte, Gunnar Mikalsen, 2014, Nomenclature and taxonomy of Telmatoscopus Eaton and Seoda Enderlein; with a discussion of parameral evolution in Paramormiini and Pericomaini (Diptera: Psychodidae, Psychodinae), Zootaxa 3878 (4), pp. 390-400 : 392-394

Telmatoscopus advena ( Eaton, 1893)

Pericoma advena Eaton, 1893: 127

= Panimerus havelkai Wagner, 1975: 1 , syn.nov.? = Telmatoscopus seguyi Vaillant, 1990: 378 . syn. nov.

Material examined. Holotype male for Panimerus havelkai Wagner, 1975 . GERMANY: Hessen, Schlitz, light trap, 23.-28.V.1974 (50.669609 ° N, 9.568590 ° E)

GERMANY: Rheinland-Pfalz, Winnweiler, Imsbach, Weisse Grube (N 49 ° 35'32.3'' E 7 ° 54' 0.7''), 355 m. a.s.l., 30.IV.2007, D. Weber leg. (RW). Hessen, Schlitz area, Rimmelswiesen (N ° 50.698860, E ° 9.484820), 2.VI.1975, R. Wagner leg. (RW). Bayern, Schweinfurt area, Oberes Garstadter Holz (N ° 49.978984, E ° 10.179700)

GREAT BRITAIN: Devon, Branscombe, 15.V.1901, A.E. Eaton leg. ( BMNH)

NORWAY: Rogaland, Finnøy, Helgøy, 5.VI-5.VII.2011. 1 m, T. Jonassen leg.; Hordaland, Bergen, Milde, Hatlehaugen. 25.V.2011. 1 m, B. Røttingen & A. Schrøder-Nielsen leg.; Sogn og Fjordane, Aurland, Fretheim, 12.VI. 1939, 1 m, Knaben leg. ( ZMUB)

Description. Adult male (n=8). Head (fig. 1A) broader than long; vertex length approximately one fifth that of head, with short neck; postocular setae difficult to distinguish from other vestiture on head, reaching half the level between side and median of head; eyebridge comprised of four facet rows, separated by half a facet diameter; interocular suture broadly U-shaped; frontal alveolar patch shaped like a narrow triangle, reaching interocular suture; length of palpomeres (n=4) 80–105 (90), 165–180 (170), 165–180 (170), 225–255 (240); labellum bulbous, setose; scapus cylindrical, pedicel elongate globular; flagellomeres asymmetrically nodiform; asymmetry less pronounced on first flagellomere; basal flagellomeres almost as long as scapus, distalmost flagellomeres slightly shorter than basal ones; length of scapus and pedicel (n=3) 100–120 (110), 75–80 (75); length of flagellomeres 1–11 (n=4) 115–135 (120), 125–140 (130), 120–150 (135), 125–150 (135), 120–150 (135), 120–145 (130), 115–145 (130), 115–145 (130), 115–140 (125), 110–140 (125), 110–135 (120); length of flagellomeres 12–14 (n=3) 100–120 (110), 90–110 (100), 80–105 (90); Thorax without special features; dorsum with setae in three broad longitudinal stripes; anepisternum, katepimeron and scutellum setose, anepimeron bare; posterior spiracle densely setose; legs without special features; Wing (fig. 1B) length 2.31–2.73 (2.51), breadth 0,87–1,05 (0,93); C thickened near base; Sc reaching level of base of R2+3; radial and medial forks about in line with each other; CuA2 not reaching level of wing forks; jugum trapezoid; Terminalia (fig. 1C) with hypandrium narrow, setose, with broad unsclerotized stripe; gonostylus (not illustrated) elongate cylindrical, with median curve, slightly longer than gonocoxite, gonocoxite subcylindrical with pointed acuminate dorsal and ventral projections meeting at middle; parameres mostly membranous, but with pair of parameral sclerites at level of basiphallic/distiphallic joint; aedeagus with basiphallus narrow, distiphallus symmetrical, with a sclerotized joint basally and curved, sabreshaped and tubular projections distally; epandrium (not illustrated) rectangular, slightly longer than broad; with single oval aperture; subepandrial plate square with rounded edges, lateral margins with thin sclerotized margins; surstyli elongate, curved, of even width throughout length; distally with 23–29 elongate tenacula which are almost as long as surstylus; tenacula distally indistinctively fringed; proctiger with hypoproct oval, epiproct triangular; both micropilose.

Remarks. Telmatoscopus advena has been recorded from the Czech Republic, Finland, France, Germany, Great Britain, Ireland, Norway, Slovakia and Sweden ( Eaton 1893, Withers & O'Connor 1992, Ježek 1998, Withers 2004, Salmela 2005, Svensson 2009, Oboňa & Ježek 2012, Kvifte & Boumans 2014). Records from Germany have hitherto only been published as Panimerus havelkai Wagner, 1975 ; however, there appears to be no significant differences between this species and Telmatoscopus advena . The synonymy was first proposed by Wagner in Salmela (2005); however, it has not previously been formalised.

Another possible synonym is Telmatoscopus seguyi Vaillant, 1990 . Most of the characters cited in the original description to distinguish this species from T. advena fall well within the variability of T. advena ; the exceptions being the number of "transverse tergapodemes" (based on the illustrations, these are apparently the margins of the subepandrial plate) and the shape of the phallomeres (parameres of Vaillant). Both of these characters may be artifacts of specimen preparation rather than characters present in the animal itself. The observed variation in ”tergapodemes” may stem from different degrees of sclerotization/bleaching of the specimens, and the phallomere shape can appear different in specimens oriented differently on microscope slides. This synonymy should, however, be confirmed through study of the type material.

Many authors write Telmatoscopus advena as Telmatoscopus advenus , but this is invalid. The name advena is latin for "stranger" and is to be treated as a noun in apposition. Nouns do not transform according to the gender of the genus name, and the correct species epithet of T. advena is therefore " advena ".