Neostasina, Rheims & Alayón, 2016

Rheims, Cristina Anne & Alayón, Giraldo, 2016, Neostasina gen. nov., a new genus of huntsman spiders from the Neotropical region (Araneae, Sparassidae, Sparianthinae), Zootaxa 4079 (3), pp. 301-344 : 304-305

publication ID	https://doi.org/10.11646/zootaxa.4079.3.1
publication LSID	lsid:zoobank.org:pub:8657278B-81C6-4571-BE3C-2B46BF1661E6
DOI	https://doi.org/10.5281/zenodo.6079047
persistent identifier	https://treatment.plazi.org/id/F9E93952-6D43-4B1A-BE93-207A29A99596
taxon LSID	lsid:zoobank.org:act:F9E93952-6D43-4B1A-BE93-207A29A99596
treatment provided by	Plazi
scientific name	Neostasina
status	gen. nov.

Treatment

Neostasina View in CoL View at ENA gen. nov.

Etymology. The generic name is a combination of the prefix Neo, derived from the Greek “neos” for “new” meaning this genus is from the New World, and the suffix Stasina in reference to the genus Stasina , from where species are being transferred. Gender is feminine.

Type species: Stasina macleayi Bryant, 1940

Diagnosis. Within Sparianthinae the new genus is characterized by three pairs of ventral spines on tibiae I and II and one pair of ventral spines on metatarsi I and II. Species of Neostasina resemble those of Decaphora by the ventral spination of tibiae and metatarsi of legs I −II, with three and one pair respectively, but can be distinguished by the male palps with dRTA simple, without spines ( Figs 15 View FIGURES 13 − 19 , 28 View FIGURES 26 − 32 , 42 View FIGURES 40 − 45 ) (complex, with many spines in Decaphora , see Rheims & Alayón 2014, figs 17, 24, 31) and by the female epigynes with lateral lobes not touching each other ( Figs 17 View FIGURES 13 − 19 , 20 View FIGURES 20 − 25. 20 − 22 , 30 View FIGURES 26 − 32 , 37 View FIGURES 33 − 39 ) (touching each other posteriorly in Decaphora , see Rheims & Alayón 2014, figs 19, 26, 34). From Pseudosparianthis it can be distinguished by having only one pair of ventral spines on metatarsi I −II (two in Pseudosparianthis ) and from Uaiuara by having three pairs of ventral spines on tibiae I −II ( 8−9 in Uaiuara ).

Description. Total length of males 5.2−11.9, of females 5.5−16.7. Prosoma slightly longer than wide; cephalic region slightly higher than thoracic region, gradually flattening posteriorly; fovea conspicuous on posterior third of prosoma. Eight eyes arranged in two rows, the anterior row straight, the posterior procurved; AME larger than ALE and more distant from each other than from ALE; PME smaller than PLE mostly more distant from each other than from PME, but can be closer or as distant as from PLE ( Figs 1−6 View FIGURES 1 − 6 ). Clypeus low, less than AME diameter. Chelicerae longer than wide, with three promarginal teeth, the basal one smallest, and two retromarginal denticles; intermarginal denticles absent; one single strong seta at the base of fang ( Fig. 7 View FIGURES 7 − 12 ). Labium slightly longer than wide. Endites convergent, with dense scopulae on internal margin. Serrula with a single row of denticles. Sternum as long as wide, very slightly projected between coxae IV. Female pedipalps with single, pectinate claw, with 3−4 teeth ( Fig. 10 View FIGURES 7 − 12 ). Legs laterigrade, mostly 2143; 2413 and 4213 can also occur. Trochanter smooth. Metatarsi I–IV distally with dorsal trilobate membrane with median hook much smaller than lateral projections. Metatarsus IV with distal ventral preening comb ( Fig. 12 View FIGURES 7 − 12 ). Tarsi and anterior half of metatarsi scopulate. Tarsal organ capsulate with keyhole-shaped opening ( Fig. 8 View FIGURES 7 − 12 ), located dorsally at the distal end of tarsi. Trichobothria present on dorsal tibiae, metatarsi and tarsi, arranged in several rows that converge to a single row at the basal end of tarsi and metatarsi. Bothrium with crescent plate, with 7−8 grooves, projected over a smooth basal plate ( Fig. 9 View FIGURES 7 − 12 ). Tarsi with pair of pectinate claws with 5−6 very slightly curved teeth and claw tufts ( Fig. 11 View FIGURES 7 − 12 ). Spination pattern in males: femora I −III: p1-1-1, d0-1-1; r1-1-1; femur IV: p1-1-1, d0-1-1, r0-0(1)-1; tibiae I −IV: p1-0-1; d0-0-1; r1-0-1; v2-2 -2-0; Metatarsi I −II: p1-1-0, r1-1-0, v2 -0-0; metatarsus III: p1-1-1, r1-1-1, v2 -0(2)-1; metatarsus IV: p1-1-1, r1-1-1, v2- 2 -0. Palp: femur p0-0-1; d0-1-2; r0-0-0(1); tibia p1-0-0; in females: femora I −II: p1-1-1, d0-1-1, r1-1-1; femur III: p1-1-1, d0-1-1, r1-1-1(0); femur IV; p1-1-1, d0-1-1, r0-0-1; tibiae I −II: p0(1)-0-0(1),, r0(1)-0-0(1) v2-2 -2-0; tibia III: p0(1)-0-0(1), d0-0-0(1); r1(0)-0-0(1), v2-2 -2; tibia IV: p1-0-1, d0-0-1, r1-0-1, v2-2 -2; metatarsi I −II: p1-0(1)-0, r1-0(1)-0, v2 -0-0; metatarsus III: p1-0(1)-0(1), r1-0(1)-0(1), v 2 -0(2)-1(0); metatarsus IV: p1-1-1, r1-1-1, v2-2 -0. Palp: femur p0-0-1; d0-0-2; patella p1, r1; tibia p2-1-0; d0-0-1; r1-1-0; metatarsus p2-1-0; r2-1-0. Opisthosoma oval, longer than wide. Male epiandrous spigots absent. Six spinnerets: anterior lateral spinnerets contiguous, conical and bi-segmented. Basal segment elongate and cylindrical, distal segment short and truncated. Posterior median spinnerets conical and short. Posterior lateral spinnerets conical and bi-segmented. Basal segment elongate and cylindrical, distal segment short and truncated. Male palp: tibia slightly shorter than cymbium; VTA situated medially ( Figs 27 View FIGURES 26 − 32 , 69 View FIGURES 68 − 71 , 90 View FIGURES 89 − 95 ) or slightly displaced retrolaterally ( Figs 50 View FIGURES 46 − 52. 46 − 48 , 107 View FIGURES 106 − 112 , 137 View FIGURES 136 − 142 ); RTA arising subdistally from tibia with dRTA single ( Figs 15 View FIGURES 13 − 19 , 55 View FIGURES 53 − 56 , 91 View FIGURES 89 − 95 ) or bifid ( Figs 74 View FIGURES 72 − 78 , 134 View FIGURES 129 − 135. 129 − 131 ) of variable shape and vRTA single ( Figs 28 View FIGURES 26 − 32 , 55 View FIGURES 53 − 56 , 66 View FIGURES 64 − 67 ) or bifid ( Figs 42 View FIGURES 40 − 45 , 70 View FIGURES 68 − 71 , 115 View FIGURES 113 − 119 ) of variable shape (absent in N. taino sp. nov., Fig. 137 View FIGURES 136 − 142 ); eRTA absent ( Figs 27 View FIGURES 26 − 32 , 51 View FIGURES 46 − 52. 46 − 48 , 65 View FIGURES 64 − 67 ) or present ( Figs 15 View FIGURES 13 − 19 , 42 View FIGURES 40 − 45 , 90 View FIGURES 89 − 95 ) with variable shape; cymbium with pronounced retrolateral rim, large round alveolus and dorsal scopulae ( Figs 15 View FIGURES 13 − 19 , 35 View FIGURES 33 − 39 , 134 View FIGURES 129 − 135. 129 − 131 ); subtegulum ring-shaped with prolateral annelli; tegulum smooth ( Figs 14 View FIGURES 13 − 19 , 34 View FIGURES 33 − 39 , 54 View FIGURES 53 − 56 ) or with protrusion, the latter arising between 3 and 7 o’clock positions ( Figs 65 View FIGURES 64 − 67 , 69 View FIGURES 68 − 71 , 90 View FIGURES 89 − 95 ); median apophysis arising from tegulum between 2 and 4 o’clock positions; conductor elongate, laminar and hyaline; TBC membranous at base of conductor; embolus simple ( Figs 27 View FIGURES 26 − 32 , 54 View FIGURES 53 − 56 , 99 View FIGURES 99 − 105 ), with dorsal keels ( Figs 67 View FIGURES 64 − 67 , 71 View FIGURES 68 − 71 , 85 View FIGURES 82 − 88 , 116 View FIGURES 113 − 119 ) or bifid ( Fig. 139 View FIGURES 136 − 142 ), arising from tegulum between 6 and 9 o’clock positions; TBE single ( Figs 41 View FIGURES 40 − 45 , 60 View FIGURES 57 − 63 , 102 View FIGURES 99 − 105 ) or bifid with main branch longer than secondary branch ( Figs 16 View FIGURES 13 − 19 , 75 View FIGURES 72 − 78 , 124 View FIGURES 123 − 128 ). Female epigyne: divided into lateral lobes and median septum; epigynal field usually longer than wide; anterior rim continuous ( Figs 43 View FIGURES 40 − 45 , 93 View FIGURES 89 − 95 , 110 View FIGURES 106 − 112 ) or divided ( Figs 17 View FIGURES 13 − 19 , 30 View FIGURES 26 − 32 , 86 View FIGURES 82 − 88 ); median septum of variable shapes, smooth ( Figs 23 View FIGURES 20 − 25. 20 − 22 , 61 View FIGURES 57 − 63 , 110 View FIGURES 106 − 112 ), with scape ( Fig. 43 View FIGURES 40 − 45 ) or protrusions (79, 96); lateral lobes with posterior triangular projections ( Figs 20 View FIGURES 20 − 25. 20 − 22 , 129 View FIGURES 129 − 135. 129 − 131 , 143 View FIGURES 143 − 148. 143 − 145 ). Female vulva: copulatory ducts short; glandular projections short, rounded with wide base ( Figs 24 View FIGURES 20 − 25. 20 − 22 , 31 View FIGURES 26 − 32 , 130 View FIGURES 129 − 135. 129 − 131 ), or elongate from two ( Figs 38 View FIGURES 33 − 39 , 77 View FIGURES 72 − 78 , 127 View FIGURES 123 − 128 ) to more than four times longer than wide ( Fig. 104 View FIGURES 99 − 105 ) or rounded on short stalks ( Figs 62 View FIGURES 57 − 63 , 147 View FIGURES 143 − 148. 143 − 145 ); spermathecae of variable shapes ( Figs 21 View FIGURES 20 − 25. 20 − 22 , 94 View FIGURES 89 − 95 , 127 View FIGURES 123 − 128 ) can be packed within a sclerotized structure, the latter with internal ducts visible by transparency ( Figs 18 View FIGURES 13 − 19 , 38 View FIGURES 33 − 39 , 44 View FIGURES 40 − 45 ) or inconspicuous ( Figs 31 View FIGURES 26 − 32 , 87 View FIGURES 82 − 88 , 121 View FIGURES 120 − 122 ); fertilization ducts slender, medially looped.