Elacatis

Pollock, Darren A., 2018, Review of the Nearctic (north of Mexico) species of Elacatis Pascoe (Coleoptera: Salpingidae: Othniinae), Zootaxa 4420 (3), pp. 301-333 : 303-304

Key to Nearctic (north of Mexico) species of Elacatis

1. Pronotal disc flat to slightly concave; elytra unicolorous red-brown, without lighter color or contrastingly lighter setal pattern ( Fig. 2 )....................................................................... E. immaculatus (Champion)

- Pronotal disc variously convex; elytra not unicolorous, with contrasting light and dark areas.......................... 2

2. Head and pronotal punctation relatively shallow and sparse, with distinct spaces around punctures (up to about the diameter of a single puncture); elytra ( Fig. 1 ) dark with 3 well-delimited, lighter areas: several irregular spots at base, transverse band near midlength, widest at suture, not reaching lateral margin, and isolated round subapical spot; legs yellow, distinctly contrasting dark dorsal body color................................................................. .. E. fasciatus (Bland)

- Head and pronotal punctation deep and dense, spaces (if any) among punctures less than diameter of single puncture; elytra dark with lighter areas, more diffuse, not as described above; legs darker, similar in color to dark dorsal body color........ 3 notal punctation rugose, borders of punctures confluent, forming contiguous, longitudinal meshes; elytra with extensive lightcolored areas ( Fig. 3 ), in approximate X-shape in most specimens, shoulders yellow.......... .. E. senecionis (Champion) - Antennae longer, reaching posterior margin of pronotum or beyond; pronotum less convex, somewhat flattened at midline; pronotal punctation less rugose, borders of punctures not forming longitudinal meshes; elytral color not as described above......................................................................................................4

4. Head, pronotum, and in some specimens, also elytra, with slight to distinct aeneous or bronzy metallic luster; elytra dark ( Figs 4–5 ) with variously developed light “zig-zag” markings (in some specimens (e.g. Fig. 6 ), elytra with greater area of lighter than darker color), but not consisting at least in part of isolated circular spots; antennae of both sexes relatively short, extending only to near posterior margin of pronotum ( Figs 4–5 ); associated with conifers in western North America............................................................................................. .. E. umbrosus (LeConte)

- Head, pronotum and elytra without metallic reflection; elytra dark, with light markings consisting in part of at least several isolated circular spots, especially on posterior half of elytra; antennae sexually dimorphic, relatively short in female (no paired pits on mentum), extending to near posterior margin of pronotum; antennae longer in males (paired pits on mentum), extending beyond hind margin of pronotum; not known to be associated with conifers....................................... 5

5. Lateral pronotal carina of most specimens with 3 tubercles (hind angle, widest point, and one in between) ( Figs 11–14 ); lateral pronotal margins evenly arcuate, anterior margin width subequal to posterior margin width; male tegmen relatively elongate ( Figs 32–33 )......................................................................................... 6

- Lateral pronotal carinae ( Figs 7–10 ) relatively smooth, tubercles at hind angles and in some specimens, slight suggestion of tubercle at maximum width; lateral pronotal margins only slightly arcuate, anterior margin width greater than posterior margin width; male tegmen relatively short, wide ( Fig. 31 )............................................ E. longicornis Horn

6. Tegmen with relatively long, narrow basal piece (length of apicale <1.5 x length basale); length of basale subequal to width ( Fig. 32 ); distribution ( Fig. 38 ): southern Great Plains (CO, KS, NE, MT)....................... E. larsoni , new species

- Tegmen with relatively short, wide basal piece (length of apicale> 1.5 x length basale); length of basale distinctly greater than width ( Fig. 33 ); distribution ( Fig. 39 ): southeastern Arizona and Mountains and Basins region of Texas ................................................................................................... E. stephani , new species

Figures

FIGURES 1–6. Dorsal habitus of Elacatis spp. 1. Elacatis fasciatus (Bland) [“Va.”], female, TL = 5.2 mm; 2. Elacatis immaculatus (Champion) [AZ: Pinal Co.], male, TL = 4.2 mm; 3. Elacatis senecionis (Champion) [TX: La Salle Co.], male, TL = 5.1 mm; 4. Elacatis umbrosus (LeConte) dark form [AZ: Coconino Co.], male, TL = 5.5 mm; 5. Elacatis umbrosus dark form [AZ: Coconino Co.], female, TL = 5.6 mm; 6. Elacatis umbrosus light form [CA: San Bernardino Co.], male, TL = 4.8 mm.

FIGURES 7–12. Dorsal habitus of Elacatis spp. 7. Elacatis longicornis Horn [CA: Riverside Co.], male, TL = 4.3 mm; 8. Elacatis longicornis [CA: San Bernardino Co.], female, TL 5.1 mm; 9. Elacatis longicornis [CA: Riverside Co.], male, TL = 3.7 mm; 10. Elacatis longicornis [CA: Riverside Co.], female, TL = 4.2 mm; 11. Elacatis larsoni, n. sp. [NE: Box Butte Co.], male holotype, TL = 5.0 mm. 12. Elacatis larsoni, n. sp. [KS: Rooks Co.], female allotype, TL = 4.4 mm.

FIGURES 13–14. Dorsal habitus of Elacatis stephani, n. sp. 13. Male holotype [AZ: Cochise Co.], TL = 5.1 mm; 14. Female allotype [AZ: Cochise Co.], TL = 5.0 mm.

FIGURES 28–33. Tegmen of male genitalia of Elacatis spp. (note: median lobe damaged/distorted in most specimens). 28. Elacatis immaculatus (Champion); 29. Elacatis senecionis (Champion); 30. E. umbrosus (LeConte); 31. Elacatis longicornis Horn; 32. Elacatis larsoni, n. sp.; 33. Elacatis stephani, n. sp.

FIGURES 38–39. 38. Distribution of Elacatis larsoni, n. sp.; 39. Distribution of Elacatis stephani, n. sp. Both maps created using www.gpsvisualizer.com.