Telosticta bidayuh, Dow & Orr, 2012

Dow, R. A. & Orr, A. G., 2012, Telosticta, A New Damselfly Genus From Borneo And Palawan (Odonata: Zygoptera: Platystictidae), Raffles Bulletin of Zoology 60 (2), pp. 361-397 : 367-372

publication ID	https://doi.org/ 10.5281/zenodo.5348617
persistent identifier	https://treatment.plazi.org/id/03DD87E5-FFE8-6159-FEE8-FCD6FE1D9856
treatment provided by	Tatiana
scientific name	Telosticta bidayuh
status	sp. nov.

Treatment

Telosticta bidayuh View in CoL , new species

( Figs. 3A View Fig , 4A View Fig , 5A, G View Fig , 6A View Fig , 7A View Fig , 8A View Fig , 19 View Fig )

Protosticta feronia View in CoL – Lieftinck, 1954: 31 (part); Orr, 2003: 38, 69 (part);? Donnelly, 1997: 124, 126

Protosticta sp. cf. feronia View in CoL – Dow & Reels, 2010: 15, 18

Material examined. — Holotype: male (SAR06_PST16), Malaysia, Sarawak, Kuching division, near Indonesian border, Annah Rais, small high gradient stream in highly disturbed forest, 26 Jan.2006, coll. RAD, to be deposited in RMNH.

Paratypes (all from Sarawak, all from Kuching Division and all currently in coll. RAD unless otherwise noted, examples to be deposited in BMNH and ZRC): male (SAR06_PST97), location as holotype, 26 Jan.2006; 2 males (SAR07_8_PST245–246) , Gunung Singgai , small forest stream, ca 250 m, 2 Oct.2008; 4 males (SAR09_10_PST365, 378–380) , Kubah National Park , stream at ca 650–700 m on Gunung Serapi, 3 Jun.2010 ; male (SAR11_12_ PST138), same location, 3 Sep.2011 ; male (SAR05_PST79), same national park, small streams on Gunung Serapi below 500 m, 13 Apr.2005 ; female (SAR07_8_PST348), same location, 3 Oct.2008; 2 males (SAR09_10_PST39–40) , female (SAR09_10_PST41), same location, 13 Jul.2009 ; male (SAR09_10_PST54), same national park, small stream near Main Trail , 28 Oct.2009 ; female (SAR06_PST9), same national park, Waterfall Trail , apparently ovipositing into the mid-rib of a leaf growing from a dripping wet cliff face, 14 Apr.2006 ; male (in ethanol, RMNH _INS_500006), same location, 21 Aug.2008 ; male, female (both teneral, SAR09 _10_PST52–53), same area, main stream above waterfall, 16 Sep.2009 ; male (SAR07_8_PST115), Kubah National Park , Belian Trail, small high gradient forest stream, ca 80 m, 29 Nov.2007 ; male (SAR07_8_PST302), same location, 15 Sep.2008; 2 males (SAR09_10_PST389–390) , same location, 25 Apr.2010 ; male (SAR11_12_PST137), Semenggoh Nature Reserve , small stream in arboretum, 7 Oct.2011 ; male (JvT 26584), Samarahan division, Serian area, Tapuh, 4–9 Jul.1958, coll. T.C. Maa , in RMNH; male (JvT 28069), no location information apart from “ Sarawak ”, 30 Sep.1950, coll. M. A. Lieftinck , in RMNH.

Diagnosis. — The male of T. bidayuh is distinguished from all other species of Telosticta except T. feronia by the combination of moderate length antehumeral stripes and long lateral pronotal processes and superior appendages with poorly developed dorsal projections and a flattened but not strongly folded apical part with at most a small notch in the tip as seen in normal lateral view. It is distinguished from T. feronia by the markings of the labrum and terminal abdominal segments and by the position of the internal projection of the superior anal appendage, placed at the point where the basal part of the appendage contracts in T. feronia but well distal to this point in T. bidayuh .

Etymology. — bidayuh , a noun in apposition. Named for the Bidayuh people; the known range of the species falls within the Bidayuh heartland.

Description of holotype male. — Head: Labium pale. Basal 3/4 of labrum blue, black along free margin. Anteclypeus blue, postclypeus shining black. Mandible bases blue in corner by clypeus, black below. Vertex and frons bronzy black, occiput shining black. Ratio of width of compound eye to width of vertex measured at level of lateral ocelli slightly over 9/10. Transverse occipital carina with lateral extremities angulated and prominent. Ocelli yellowish. Antenna with scape pale yellowish, remainder missing.

Thorax: Prothorax, largely pale cream except to rear of propleuron where there are irregular dark markings, a few brown patches at rear of middle pronotal lobe and whole posterior pronotal lobe, where black, becoming dark greyish white on lateral process, which is of moderate length ( Fig. 3A View Fig ) with the tip well above the level of the lower margin of the propleuron. Synthorax: Mesepisternum bronzy black, with a pair of blue antehumeral stripes, occupying ca 2/3 of the length ( Fig. 4A View Fig ). Antealar triangles blue. Mesepimeron bronzy black. Metepisternum largely occupied by a pale band, becoming blue towards wing bases, with a bronzy black wedge below, this extending from the wing bases to just before the spiracle. Metepimeron almost entirely pale. Venter of synthorax pale. Legs: each with coxa and trochanter cream, femur cream with dark stripe on extensor surface and some obscure dark markings at side, with obscure grey markings, black above tibia. Tibia black immediately below femur, then pale, black just above tarsus, which is pale with some black areas and brown claws. Wings: 13 Px in Fw, 12 Px in Hw. Vein ab absent. Arculus slightly distal to Ax2. R 4 arising at or distal to subnodus, IR 3 joined to it by a short stalk. Pterostigma with costal side slightly shorter than anal side, dark brown with white border, covering slightly more than one underlying cell.

Abdomen: Largely pale brown, darkening to rear. S1 whitish, brown behind posterior carina and narrowly before it. S2 with a pale basal annulus, just divided dorsally, laterally this extending to the posterior carina as a pale wedge, otherwise brown. S3–6 brown above, pale laterally with a well defined boundary between the two shades on S3–4, becoming less clear after this. A narrow pale basal annulus, narrowly interrupted dorsally on S3 but complete on S4–6, this continued narrowly as a pale stripe along the lower margin of each segment, becoming indistinct apically. S7 similar but black dorsally, with the basal annulus broadly interrupted dorsally. S8–10 black with a pale blue dorsal bilobed marking on S8, this extending laterally at the segment base, becoming cream coloured halfway down the side and continued narrowly along the lower margin of the tergite for the length of the segment ( Fig. 5A, G View Fig ). Anal appendages ( Figs. 6A View Fig , 7A View Fig , 8A View Fig ) largely brown. Superior appendage slightly less than 2.5 times the length of S10, with interior projection at slightly less than half length, directed inwards at base so just visible in lateral view and then upwards with a sharp tip. Dorsal projection a small but well defined rounded knob directed inwards and slightly upwards ( Fig. 6A View Fig ). Expanded dorsal-ventrally in distal 1/3, with a notch at tip, difficult to see in normal lateral view. Inferior appendage ca same length as superior, scoop only slightly expanded, in ventral view turning rather abruptly inwards and then straightening in terminal part, where concave surface up and slightly outturned, the spine large and directed inwards, upwards and to the rear ( Fig. 8A View Fig ).

Measurements (mm): abdomen without anal appendages 33.5, superior appendage ca 1.25, Hw 18.

Description of female paratype. — Based on (SAR06_PST9); as male except as noted.

Head: blue on labrum with a narrow triangular excision centrally.

Thorax: lateral process of posterior pronotal lobe less pale than in male.

Wings: R 4 arising distal to subnodus in all wings. 12 Px in Fw, 10–11 Px in Hw. Pt brown.

Abdomen: pale basal annulus on S7 broad, not interrupted dorsally. S8 mostly pale blue, S9 dark brown with obscure paler areas basally and laterally. Superior anal appendages shorter than S10. Ovipositor mottled dark brown and cream, long, valves extending well beyond tips of superior anal appendages.

Measurements (mm): Abdomen without anal appendages or ovipositor 31, Hw 19, ovipositor slightly less than 2.

Variation in paratypes. — Males: there is some slight variation in the length of the antehumeral stripes, and in the

shape of the lateral processes of the pronotal posterior lobe. In some individuals the pale mark on the dorsum of S8 is not bi-lobed, in the specimen collected by Lieftinck without a specific location there are a pair of tiny widely spaced basal dorsal lateral spots on S9. The notch in the tip of the superior appendage appears slightly deeper and/or wider in a few specimens, but to a large extent this is due to the way that the appendages are laying in the individual specimen.

Females: Little variation except in size is apparent, but in one individual from Kubah there is a pair of tiny blue dorsal spots on S9.

Measurements (mm): Males: Abdomen without anal appendages 31–36.5, Hw 17.5–20.5. Wings with 11–13 Px in Fw, 10–12 in Hw. Females: Abdomen without appendages or ovipositor 31–33, Hw 19–21. Wings with 12–13 Px in Fw, 10–11 in Hw.

Biological notes. — This species is found at high gradient forest streams, usually small but teneral individuals have been found beside a larger torrential stream at Kubah National Park.At Kubah National Park it occurs together with T. serapi at many locations, but typically occurs at lower densities.

Females have been found perched on vegetation at wet cliff faces beside waterfalls on several occasions.

Distribution. — Known from Sarawak west of the Lupar River, with records from Serian in Samarahan division, and in Kuching division from the Matang Range (including Kubah National Park), Semenggoh Nature Reserve, and Annah Rais ( Fig. 19 View Fig ). Although the specimen collected by Lieftinck has no location information with it other than “ Sarawak ”, Lieftinck’s 1950 visit to the state was confined to Sarawak west of the Lupar and his specimen probably comes from somewhere close to Kuching.