Clinopodes, C. L. Koch, 1847

Bonato, Lucio, Iorio, Étienne & Minelli, Alessandro, 2011, The centipede genus Clinopodes C. L. Koch, 1847 (Chilopoda, Geophilomorpha, Geophilidae): reassessment of species diversity and distribution, with a new species from the Maritime Alps (France), Zoosystema 33 (2), pp. 175-205 : 196-197

KEY TO THE SPECIES OF CLINOPODES

1. All or almost all openings of the coxal organs distinctly grouped into 2-4 clusters on each side, completely or almost completely covered by the metasternite ( Fig. 1E ) .............. 2

— Openings of the coxal organs scattered or only partially grouped close to the metasternite, and not covered by the latter ( Fig. 1F ) ........................................................................ 6

2. No coxal pores isolated from the clusters .................................................................... 3

— One small pore on each coxopleuron, isolated from the clusters ( Fig. 1E ) .................. 4

3. The largest sternal pore-fields on the posterior leg-bearing segments extending forward to reach or even extending beyond the mid-length of the metasternites ( Fig. 1C ) .......... ................................................................................................................. C. escherichii

— The sternal pore-fields on the posterior leg-bearing segments limited to 1/3-2/5 of the length of the metasternites ( Fig. 1D ) ............................................................ C. flavidus

4. The largest sternal pore-fields on the posterior leg-bearing segments extending forward to reach the mid-length of the metasternites ( Fig. 1C ) ............................ C. intermedius

— The sternal pore-fields on the posterior leg-bearing segments limited to 1/3 of the length of the metasternites ( Fig. 1D ) ..................................................................................... 5

5. Anterior denticles of the forcipular coxosternite not very prominent, usually distinctly wider than long ( Fig. 1A ). Usually more than 65 leg-bearing segments ......................... ................................................................................................... C. caucasicus n. comb.

— Anterior denticles of the forcipular coxosternite very prominent, usually only slightly wider than long or as wide as long ( Fig. 1B ). Usually less than 65 leg-bearing segments ...................................................................................................... C. verhoeffi n. nom.

6. Chitin-lines reaching the condyles of the forcipular coxosternite ( Fig. 1B ) ................. 7

— Chitin-lines not reaching the condyles of the forcipular coxosternite ( Fig. 1A ) ........... 8

7. Anterior denticles of the forcipular coxosternite not very prominent, wider than long ( Fig. 1A ) .................................................................................... C. latisternus n. comb.

— Anterior denticles of the forcipular coxosternite not very prominent, about as long as wide ( Fig. 1B ) ................................................................................................... C. rodnaensis

8. The sternal pore-fields on the posterior leg-bearing segments limited to 1/3 of the length of the metasternites ( Fig. 1D ). No coxal pore distinctly isolated from the other pores ... .................................................................................................................. C. skopljensis

— The largest sternal pore-fields on the posterior leg-bearing segments extending forward to reach or even extending beyond the mid-length of the metasternites ( Fig. 1C ). One coxal pore on each coxopleuron, isolated from all other pores ( Fig. 1E, F ). .......................... 9

9. Isolated coxal pore larger and posterior to all other pores on each coxopleuron ( Fig. 1F ) .............................................................................................................. C. carinthiacus

— Isolated coxal pore smaller and lateral to the other pores on each coxopleuron ( Figs 1E ; 3E ) ........................................................................................................ C. vesubiensis n. sp.

Figures

FIG. 1. — Main differential characters between species of Clinopodes C. L. Koch (line drawings from representative specimens, ventral view): A, forcipular segment of C. carinthiacus (Latzel, 1880) (♀, 33 mm long, from Cansiglio, Italy); B, forcipular segment of C. rodnaensis (Verhoeff, 1938) (♀, 32 mm long, from Casa de Piatra, Romania); C, metasternites of the most posterior leg-bearing segments of C. escherichii (Verhoeff, 1896) (♀, 55 mm long, from Boltoz Dağ, Turkey); D, metasternites of the most posterior legbearing segments of C. verhoeffi n. nom. (♀, 35 mm long, from Casa de Piatra, Romania); E, metasternite and coxopleura of ultimate leg-bearing segment of C. caucasicus (Selivanov,1884) n. comb.(♀, 55 mm long, from Tbilisi,Georgia);F, metasternite and coxopleura of ultimate leg-bearing segment of C. carinthiacus (Latzel, 1880) (♀, 33 mm long, from Cansiglio, Italy). Abbreviations: c, condyle; cl, chitin-line; d, anterior denticle of forcipular coxosternite; m, metasternite of ultimate leg-bearing segment; p, isolated coxal pore; pf, pore-field. Scale bars: 0.2 mm.

FIG. 3. — Clinopodes vesubiensis n. sp. (line drawings of the holotype, ♀, 38 mm long, from Lucéram, France): A, anterior part of cephalic capsule, without maxillae and mandibles, ventral view, areolation only partially drawn; B, left half of the maxillary complex, ventral view;C, forcipular segment,ventral view;D, leg-bearing segment 12,without legs,ventral view;E, ultimate leg-bearing segment and postpedal segments, ventral view. Scale bars: 0.2 mm.