Andecladodes ovalis, Bocakova & Campello-Gonçalves & Silveira, 2022

Bocakova, Milada, Campello-Gonçalves, Lucas & Silveira, Luiz Felipe Lima Da, 2022, Phylogeny of the new subfamily Cladodinae: neotenic fireflies from the Neotropics (Coleoptera: Lampyridae), Zoological Journal of the Linnean Society 195, pp. 1181-1199 : 1196-1198

publication ID

https://doi.org/ 10.1093/zoolinnean/zlab091

publication LSID

lsid:zoobank.org:pub:73D99D73-1735-4F27-844E-796874E4A040

DOI

https://doi.org/10.5281/zenodo.6986028

persistent identifier

https://treatment.plazi.org/id/03DB87C0-FFE7-885D-F4A6-6F7B11EB7D42

treatment provided by

Plazi

scientific name

Andecladodes ovalis
status

sp. nov.

ANDECLADODES OVALIS SP. NOV.

( FIGS 3E View Figure 3 , 4E View Figure4 , 5E View Figure 5 , 6C View Figure 6 , 7D View Figure 7 , 8G, H View Figure 8 , 9D, 10F1, F2)

Z o o b a n k r e g i s t r a t i o n: h t t p: / / z o o b a n k. o r g / N o m e n c l a t u r a l A c t s /0 1 6a5e e 0-e b f8- 4 5b e - b3 4 0- 65b3c6d39192

Holotype: Male , ‘ ECUADOR, Baron’, without other data ( MNHN).

Diagnosis: Body oval, widest in elytral midlength, basal half of elytra laterally yellow ( Fig. 3E View Figure 3 ), antennae with slender lamellae, their length decreasing distally, posterior corners of pronotum obtuse.

Description: Body dark brown, only basal half of each elytron laterally yellow and luminous spots on sternum VIII and pygidium pale. Head with small eyes, interocular distance 1.6× longer than eye diameter. Antennae short, reaching humeral areas. Antennomere I stout, triangular, with apex prolonged in a short blunt horn, as long as half of antennomere width. Antennomeres II and III small, lamellae long, slender. Length of antennomeres III–X increasing to apex, while length of their lamellae decreasing apically (lamella of antennomere III the longest). Mandibles stout, arcuate to falcate ( Fig. 6C View Figure 6 ), labrum connate to frontoclypeus, anterior margin weakened, clypeolabral suture inconspicuous. Maxillary palps with terminal palpomere broad, about 1.5× longer than subapical palpomere, more or less compressed, labial palps with terminal palpomere elongate, narrowed apically, both palps setose. Pronotum with anterior margin almost semicircular, lateral margins moderately divergent backwards ( Fig. 3D View Figure 3 ), discal area elevated, finely pubescent, anterior and lateral edges of pronotum with broad punctures, posterior corners obtuse. Scutellum triangular. Elytra broadly oval, about 1.8× longer than humeral width, widest in elytral half. Longitudinal costae largely obsolete, costa 1 absent in distal half, costa 2 fully developed, costae 3 and 4 indistinct, marginal costa present, as strong as costa 2, sutural margins dehiscent. Pygidium widened apically, median extension as long as the lateral ( Fig. 8H View Figure 8 ). Median point of sternum VIII broad ( Fig. 8G View Figure 8 ), sternum IX emarginated distally ( Fig. 9D View Figure 9 ). Syntergite asymmetrical. Phallus broadly widened in basal third, gently extended in distal onefifth ( Fig. 10F View Figure 10 1–F View Figure 1 2 View Figure 2 ), apex inconspicuously cleft. Body length: 16 mm, humeral width: 6.7 mm, width in elytral midlength: 7.6 mm. Female unknown.

Etymology: Named in reference to the body outline.

Distribution: Ecuador.

LIST OF GENERA OF CLADODINAE

Cladodes Solier, 1849

= Rhipidophorus Solier, 1849

Ledocas Olivier, 1885

KEY TO GENERA OF CLADODINAE 1. Flabellum of antennomere III markedly shorter than IV, antennomeres III–X of same length or increasing in length up to midlength .................................................................................................................................2

– Flabellum of antennomere III the longest of all, length of lamellae of antennomeres III–X gradually decreasing towards apex, length of antennomeres III–X increasing distally .................6 ( Dodacles View in CoL lineage)

2. Outline parallel to slightly elliptic, elytral marginal costa facing downwards in ventral view ...................3

– Outline broadly elliptic, marginal costa facing outwards in ventral view.....................................................5

3. Clypeus connate to frons, pronotum anteriorly rounded ................................................................................4

– Clypeus connected to frons by a membrane, pronotum almost triangular................................. Cladodes View in CoL s.s.

4. Flabellum of antennomere III up to half as long that of IV, syntergite with paired, horn-like appendages, phallus with paired, longitudinal dorsal keels ........................................................................... Cladodes ater View in CoL

– Flabellum of antennomere III slightly shorter than IV, syntergite lacking paired, horn-like appendages, phallus without paired, longitudinal dorsal keels................................................................................ Ledocas View in CoL

5. Clypeus entirely connate to frons, anterolateral region of pronotum with well-delimited punctures of similar size (not forming rudimentary vitreous spots), phallus lacking paired, longitudinal dorsal keels .............................................................................................................................................. Cladodes gloriosus View in CoL

– Clypeus usually connected to frons by membrane; if connate, just at central third, anterolateral region of pronotum with punctures of different sizes that often fuse to form rudimentary vitreous spots, phallus with paired, longitudinal dorsal keels ............................................................................................ Nyctocrepis

6. Scape with inner margin acuminate, often forming a well-defined spine, mandibles arcuate.....................7

– Scape rounded, mandibles right-angled or almost so .....................................................................................8

7. Elytral inner margin straight to slightly dehiscent, pygidium with posterior corners longer and wider than median, sometimes with prosternum fused to hypomera, phallus longer than paramere .................................................................................................................................. Andecladodes gen. nov.

– Elytral inner margin moderate to strongly dehiscent, pygidium with posterior corners shorter and narrower than median, prosternum never fused to hypomera, phallus always shorter than paramere .... Dryptelytra View in CoL

8. Pygidium transverse, posterior corners feebly developed and rounded, elytron never dehiscent ................................................................................................................................... Brasilocladodes gen. nov.

– Pygidium as long as wide, posterior corners well developed and acute, elytron slightly to strongly dehiscent .............................................................................................................................................................. Dodacles View in CoL

Nyctocrepis Motschulsky, 1853

subgenus Fenestratocladodes Pic, 1935

Dodacles Olivier, 1885

Brasilocladodes gen. nov.

Dryptelytra Laporte, 1833

Andecladodes gen. nov.

MNHN

Museum National d'Histoire Naturelle

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Lampyridae

Genus

Andecladodes

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