Ganelius zombi Paulsen, 2017

Paulsen, M. J., 2017, Revision of the endemic Madagascan stag beetle genus Ganelius Benesh, and description of a new, related genus (Coleoptera: Lucanidae: Lucaninae: Figulini), Insecta Mundi 2017 (592), pp. 1-16 : 8-11

Ganelius zombi Paulsen , new species

Type material. Holotype male ( Fig. 13 ), CASC, labeled ( Fig. 15 ): a) “ MADAGASCAR: Toliara Prov / Parc National de Zombitse / 19.8 km 84° E Sakaraha / elev 770 m 5–9 Feb 2003 / 22° 50′ 36″ S 044° 42′ 36″ E ”; b) “coll. Fisher, Griswold et al. / California Acad. of Sciences / general collecting, tropical dry / forest, coll. code: BLF7509 ”; c) “Collection of the / California Academy of Sciences, San / Francisco, Calif.”; d) on red paper, “ Ganelius / zombi ♂ / HOLOTYPE / des. M.J. Paulsen 2017”.

Three female paratypes, 2 CASC, 1 MJPC, labeled: a–c) as holotype. One male, one female paratypes, CASC, labeled: a) “MADAGASCAR: Mahajanga / Prov: Parc National Tsingy / de Bemaraha, 3.4 km 93° E / Bekopaka, Tombeau Vazimba / Elev 50 m 6–10 Nov. 2001 / 19° 8′ 31″ S 44° 49′ 41″ E ”; b) “coll: Fisher, Griswold et al. / California Acad. of Sciences / general collecting, tropical dry / forest, coll. code: BLF4231”; c) as holotype. One female paratype ( CASC), labeled: a) MADAGASCAR: Mahajanga / Prov: Parc National Tsingy / de Bemaraha, 2.5 km 62° ENE / Bekopaka, Ankidrodroa River / elev 100 m 11–15 Nov. 2001 ”; b) “ 19° 7′ 56″ S 44° 48′ 53″ E / coll: Fisher, Griswold et al. / California Acad. of Sciences / general collecting – tropical dry / forest on Tsingy, BLF4342”; c) as holotype. One male, one female paratypes ( CASC, MJPC) labeled: a) “MADAGASCAR: Mahajanga / Prov. Forêt de Tsimembo , / 8.7 km 336° NNW Soatana / elev 20 m 21–25 Nov. 2001 / 19° 1′ 17″ S 44° 26′ 26″ E ”; b) “coll: Fisher, Griswold et al. / California Acad. of Sciences / general collecting in tropical / dry forest code: BLF4509”; c) as holotype. One male paratype ( CASC) labeled: a) “MADAGASCAR: Mahajanga / Prov: Parc National Tsingy /de Bemaraha, 10.6 km ESE / 123° Antsalova, elev 150 m / 16–20 November, 2001 / 19° 42′ 34″ S 44° 43′ 5″ E ”; b) “coll: Fisher, Griswold et al. / California Acad. of Sciences / at light in tropical dry forest / on Tsingy, BLF4431”; c) as holotype. One paratype male ( CASC) labeled: a) handwritten, “ Nigidius / madagas- / cariensis ”; b) “Collection of the / California Academy of Sciences, San / Francisco, Calif.”. One paratype female ( MNHN) labeled: a) “Forêt de l’ / Ankarafantsika” ; b) “Museum Paris”. All paratypes labeled on yellow paper, “ Ganelius / zombi (♂ or ♀) / PARATYPE / des. M.J. Paulsen 2017” .

Description, holotype male. Length: 16.4 mm. Width: 6.6 mm. Color: Piceous black. Head: Surface velvety shagreened, impunctate basally, anterior half of frons shagreened with fine, shallow punctures, becoming shiny near margins with larger punctures laterally. Clypeus short, emarginate on each side, apex acuminate with median denticle. Clypeolabral suture obsolete. Labrum bifurcate, visible dorsally, just below clypeus. Suprantennal tubercle weakly elevated but distinct, elongate. Ocular canthus broad, rounded, laterally produced, convex; canthus not contiguous with anterior and posterior margins of head; postocular process a large, compressed, transverse plate. Antennal club moderate, short (shorter than scape), antennomeres tomentose only distally. Mandibles (to true apex) shorter than head, apex triangular, internal tooth bifurcate. Dorsal surface produced into long, vertical ramus (false apex); ramus with apex truncate, conjoined to triangular subapical tooth, margin not serrate internally, basal lobe forming an obtuse tooth remote from smaller internal tooth. Mentum broad, distinctly emarginate, surface coarsely, contiguously punctate. Pronotum: Form broad, wider than elytra. Anterior angles transversely truncate. Surface smooth with minute punctures. Anteromedial tubercle small, anterior margin produced on either side. Elytra: Form subparallel, widest at middle. Surface with serially punctate striae, striae impressed basally (rugose at base) becoming weakly impressed towards apex, striae 7–9 almost obsolete except medially, striae very with regular sides without punctures crenulating intervals; strial punctures elongate. Intervals strongly convex basally, becoming flat towards apex, punctate with fine, shallow punctures in 3–5 irregular rows. Legs: Protibia with 7 acute teeth decreasing in size proximally. Mesotibiae and metatibiae with 1 large external tooth below middle with 1–3 smaller accessory teeth proximally. Abdomen: Male genitalia ( Fig. 16 ) with permanently everted internal sac (flagellum) whip-like, 45 mm.

Paratype variation. Males (n = 4). Length: 13.0– 16.1 mm. Width: 5.2–6.9 mm. Postocular process varying from compressed as in holotype to bulbous; reduced in smaller males. Females (n = 7). Length: 13.2–18.9 mm. Width: 5.2–6.9 mm. The females ( Fig. 14 ) differ from the holotype male in the following characters: Clypeal apex with median tooth larger. Labrum lower than clypeus, not distinctly visible in dorsal view. Suprantennal tubercles more strongly elevated. Antennal club smaller. Mandibles lacking dorsal ramus, subapical dorsal tooth moderately large, obtuse. Anteromedial pronotal tubercle strongly produced in large females.

Remarks. The strongly concave ocular canthus of females of this species is immediately recognizable ( Fig. 14 ). Males are somewhat less distinctive, but the elytral striae of the disc are narrower and have straight margins that are not crenulated by the punctures as in G. madagascariensis and G. oberndorferi . Etymology. The specific epithet is a shortened form of the type locality Zombitse with which I originally labeled the specimens during the revision, and which gradually stuck. It is used as a noun in apposition based on an arbitrary (non-Latin) combination of letters.

Distribution ( Fig. 36 ). MADAGASCAR: MAHAJANGA: Forêt d’Ankarafantsika; Forêt de Tsimembo near Soatana; P.N. Tsingy de Bemaraha. TOLIARA: P.N. Zombitse.

Figures

Figures 13–16. Ganelius zombi Paulsen, n.sp. 13) Male holotype, dorsal habitus. 14) Female paratype, dorsal habitus. 15) Holotype labels. 16) Male genitalia, flagellum, ~50 mm.

Figures 33–37. Species distributions. 33) Ganelius madagascariensis. 34) G. oberndorferi. 35) G. gnamptus. 36) G. zombi. 37) Agnelius nageli.