Colistra Davies

Stiller, Michael & Webb, Michael D., 2022, Leafhoppers of the Fynbos Biome of South Africa: Colistra, Proekes, Proekoides and a new genus (Insecta, Hemiptera, Cicadellidae, Deltocephalinae, Bonaspeiini), Zootaxa 5199 (1), pp. 1-79 : 5-10

publication ID	https://doi.org/ 10.11646/zootaxa.5199.1.1
publication LSID	lsid:zoobank.org:pub:61CD7E40-DD47-4FAC-A880-42B505A84AE8
DOI	https://doi.org/10.5281/zenodo.7255473
persistent identifier	https://treatment.plazi.org/id/03D987B7-FFC4-FF8E-8FB9-DB65FA527678
treatment provided by	Plazi
scientific name	Colistra Davies
status

Treatment

Colistra Davies View in CoL

Figs 1–14 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4 View FIGURE 5 View FIGURE 6 View FIGURE 7 View FIGURE 8 View FIGURE 9 View FIGURE 10 View FIGURE 11 View FIGURE 12 View FIGURE 13 View FIGURE 14 , 31 View FIGURE 31

Colistra Davies, 1988: 215 View in CoL , 216.

Type species: Caffrolix parvulus Linnavuori, 1961 View in CoL , by original designation.

Revised generic diagnosis and description

Diagnosis.

1. Small leafhopper; male and female: length from apex of crown to apex of tegmina 2.8–3.2 mm; crown length medially 0.5–0.6 mm; crown angle 92–103°.

2. Dorsum of head in male and female with brown to dark brown color pattern with apex of crown with deltate, paired, proximate marking and circular or amorphous marking on disc, sometimes small ovoid paired making near base of head ( Figs 1A–D View FIGURE 1 , 2A–E View FIGURE 2 , 3A–Q View FIGURE 3 ), markings sometimes variable ( Fig. 3A, H, I View FIGURE 3 ), rarely immaculate ( Fig. 3K View FIGURE 3 ).

Terminalia in male:

3. Valve crescentic ( Figs 7C View FIGURE 7 , 9C View FIGURE 9 , 11C View FIGURE 11 ), length/width=0.1–0.2.

4. Pygofer deeply incised in dorsal view; pygofer lobe in lateral view, acutely triangular, mediodorsal margin with straight process, with subapical, ventral tooth; 3–6 macrosetae, rarely 8 (length 63–143 µm) ( Figs 7B, F View FIGURE 7 , 9B, D–F View FIGURE 9 , 11B, D View FIGURE 11 ).

5. Segment 10 (of anal tube) large, wide (0.6–0.7 times as wide as width across pygofer at point of greatest width of segment 10), elongate, reaching apex of pygofer lobe (dorsal view, length/width=0.9–1.1) ( Figs 7A View FIGURE 7 , 9A View FIGURE 9 , 11A View FIGURE 11 ).

6. Connective wider than long (greatest length/greatest width=0.5–0.6), stem short, wide, arms slightly shorter to much longer than stem (length arm/length stem=0.7–2.6).

7. Aedeagus with base of anterior margin of shaft edentate; either dorsal apodeme present and preatrium reduced or dorsal apodeme absent and preatrium produced.

Terminalia in female:

8. Sternite 7 posterior margin variable, either uniformly rounded to sublinear ( Figs 8A View FIGURE 8 , 10A, B View FIGURE 10 , 14D, E View FIGURE 14 ), or with deep or shallow notch flanked by rounded lobes ( Figs 12A–C View FIGURE 12 , 14A–C View FIGURE 14 ). Greatest length/greatest width=0.4–0.5, depth of notch/greatest length=0.2–0.3.

9. Valvula 3 protruding one quarter of pygofer length ( Fig. 2G View FIGURE 2 ); with 1–3 rows of marginal to submarginal setae ( Figs 8M View FIGURE 8 , 10F View FIGURE 10 , 12K View FIGURE 12 , 14H, I View FIGURE 14 ).

10. Valvifer 2 sculpture pore-like and microtrichia ( Figs 8F View FIGURE 8 , 10J View FIGURE 10 , 12G View FIGURE 12 , 14J View FIGURE 14 ).

Etymology. Retrospectively, not provided by Linnavuori (1961) or Davies (1988). Latin colis, penis, masculine and two possible suffixes denoting either tools and means; e.g., -tra, -tron, -trum, or diminutive suffixes with derogatory implication, wild resemblance -aster, -tra, -trum; (e.g., Aspidistra elatior , common aspidistra, cast-iron plants, Asparagaceae , Plantae). Gender masculine, determined by the root ( Brown 1956).

Description

Male, female and nymph.

Color.

Head. Ground color ochraceous to stramineous. Apex of crown with paired triangular marking, large ( Figs 1A–C View FIGURE 1 , 2C, E View FIGURE 2 , 3D, F, G, N, Q View FIGURE 3 ), or small ( Figs 1D View FIGURE 1 , 2D View FIGURE 2 , 3C, M View FIGURE 3 ), sometimes absent ( Fig. 3K View FIGURE 3 ). Disc with brown ( Figs 1A–C View FIGURE 1 , 2C, E View FIGURE 2 , 3C, D, N–Q View FIGURE 3 ) to light brown ( Figs 1D, I View FIGURE 1 , 2A, B View FIGURE 2 , 3B View FIGURE 3 ) circular or amorphous marking, sometimes with wide ( Figs 1B View FIGURE 1 , 2E View FIGURE 2 , 3B, 3G View FIGURE 3 ) or narrow lateral extension ( Figs 1A View FIGURE 1 , 3F, L, N–Q View FIGURE 3 ), large ( Figs 1B View FIGURE 1 , 2E View FIGURE 2 , 3D, P View FIGURE 3 ) or small ( Fig. 3E View FIGURE 3 ) or divided medially ( Figs 1D View FIGURE 1 , 2B, D View FIGURE 2 , 3H, I, M View FIGURE 3 ), or sometimes absent ( Fig. 3A, K View FIGURE 3 ). Posterior margin of head with 1–2 small ovoid markings present (e.g., Figs 1A View FIGURE 1 , 2E View FIGURE 2 , 3F, O View FIGURE 3 ) or absent or faded.

Face. Ground color ochraceous to stramineous, frontoclypeus with numerous dark brown lateral arcs ( Figs 1F View FIGURE 1 , 2H View FIGURE 2 ).

Nymph. Variable, reddish, brown or ochraceous, mottled with darker marking ( Fig. 3R–T, V View FIGURE 3 ), median pale longitudinal line ( Fig. 3R–T, V View FIGURE 3 ), or median reddish-brown line ( Fig. 2J, L, M View FIGURE 2 ) with head marked as in adult ( Fig. 3T, U, V View FIGURE 3 ) or unmarked ( Figs 2K View FIGURE 2 , 3R, S View FIGURE 3 ).

Tegmina. Brown marking in margin of outer costal, inner discal cell and inner claval cell. Cells appear opaque white due to reflectance of hind wing under tegmina; veins brown; costal cells yellow or translucent.

Hind wing. Elongate wing light brown to brown to grey-brown translucent, veins brown; short wing pale white opaque or light brown opaque, veins variable, light brown or without color.

Morphology.

Head. Crown angle 92–104°. Wider than pronotum.

Face. Postclypeus longer than wide, with lateral margins parallel to slightly convergent, anterior and posterior margins convex; posterior margin extended beyond lora and posterior margin of gena below lorum (=maxillary plate) narrow.

Ocellus. Ocellus diameter/interocular distance=0.4–0.6.

Pronotum. Lateral margin short, smooth.

Tegmina. Length/width=1.89–2.12; four apical cells, outer and central anteapical cells, inner anteapical cell and inner discal cell fused or without outer anteapical cell ( Figs 7I View FIGURE 7 , 8N View FIGURE 8 , 9O View FIGURE 9 , 10O View FIGURE 10 , 11M View FIGURE 11 , 12P View FIGURE 12 , 13Q View FIGURE 13 , 14N View FIGURE 14 ); appendix absent.

Hind wing. Shorter than tegmina, elongated with costal margin sublinear ( Figs 7J View FIGURE 7 , 8O View FIGURE 8 , 9N View FIGURE 9 , 10P View FIGURE 10 ) or short, about as wide as long with costal margin shallowly convex ( Figs 11N View FIGURE 11 , 12Q View FIGURE 12 , 13R View FIGURE 13 , 14M View FIGURE 14 ); opaque; about one third shorter and narrower respectively than tegmina length and width; length/width=1.1–2.4. Ratio of length and width of tegmina/length and width of hind wing of male and female=1.9–2.0.

Chaetotaxy. Profemur row AV 9–11 short, wide setae, row IC 7–9 long, fine setae, AV 1 single, AM 1 single. Spinulation of protibia 1+4, mesotibia 4+4, metafemur apex 2+2+1. Metatarsus 1 apical plantar setae include four rounded platellae, one acute seta (lateral) or rarely five round and one acute; metatarsus 2 apical plantar setae include one acute seta, two rounded platellae and one acute seta or rarely three round, one acute.

Measurements.

Male and female similar dimensions (n=339). Apex of crown to apex of tegmina 2.7–3.4 mm, apex of crown to apex of abdomen 2.8–3.9 mm; crown length 0.5–0.6 mm; crown length next to eye 0.3–0.4 mm; pronotum length 0.4 mm; head width 1.1–1.3 mm; pronotum width 1.0– 1.3 mm; ocellus diameter 24–31 µm; interocular distance 45–66 µm; crown angle 92–104°; crown length/crown length next to eye=1.3–1.5; head width/pronotum width=1.0–1.1; ocellus diameter/interocular distance=0.4–0.6; crown length/pronotum length=1.0–1.5; pronotum length/pronotum width=0.3–0.4.

Terminalia.

Male

Segment 10. Large, incised deeply into pygofer, almost to base of pygofer, lateral margin convergent, sublinear ( Figs 7A View FIGURE 7 , 9A View FIGURE 9 ) or convex ( Fig. 11A View FIGURE 11 ), in dorsal view length/width=0.9–1.1, in lateral view compressed ( Figs 7B View FIGURE 7 , 9B View FIGURE 9 , 11B View FIGURE 11 ). Greatest width of segment 10 relative to greatest width of pygofer at point of greatest width of segment 10=0.6–0.7. Posterior margin of anal tube extended beyond posterior margin of pygofer lobe, or as far as pygofer lobe.

Pygofer. In lateral view longer than greatest width; anterior margin oblique, anterior apodeme elongate, angled slightly ventrad (e.g., Fig. 7B View FIGURE 7 ); basolateral suture variably curvate, commonly shallowly convex, rarely strongly convex, incised less than half-way into pygofer lobe ( Fig. 11B View FIGURE 11 ) or right-angled extending about 1/2 into pygofer lobe ( Figs 7B View FIGURE 7 , 9B View FIGURE 9 ); region posteriad of basolateral suture glabrous. Pygofer lobe, in lateral view, acutely triangular, extending almost to base of pygofer; 4–5 macrosetae, rarely eight, laterad of medial area of origin of pygofer process, 55–76 µm in length. Sclerotized process with origin at dorsomedial margin, process straight, orientation posteroventrad, tooth of process ventral, subapical ( Figs 7B, F View FIGURE 7 , 9B, D, E View FIGURE 9 , 11B, D View FIGURE 11 ), mediad of membranous lobes. Apex of lobe membranous or desclerotized, weakly demarcated ( Fig. 7F View FIGURE 7 ) or well differentiated ( Figs 9D View FIGURE 9 , 11D View FIGURE 11 ) in lateral view, medially with numerous microtrichia and few short setae; folded over subapex of sclerotized process ( Fig. 9E View FIGURE 9 ). Subapex medially with ovoid, membranous structure covered with microtrichia and few short setae ( Figs 7F View FIGURE 7 , 9E View FIGURE 9 , 11D View FIGURE 11 ). Lobe with dorsal margin near origin of process with variable, recurved, tooth-like or triangular process, about as long as wide, orientation dorsal or mediolateral, or reduced, often obscured by macrosetae, viewed better dorsolaterally.

Subgenital plate. Extended beyond posterior margin of pygofer lobe and rarely beyond posterior margin of anal tube; greatest length/greatest width=0.8–1.5; medial margins contiguous or narrowly divergent distally; apex broadly rounded; lateral margin shallowly convex. Macrosetae in 2–3 rows near apex, single row medially, macrosetae absent basally, medial macrosetae about as long as median width of subgenital plate, lateral macrosetae shorter.

Valve. Crescentic ( Figs 5C View FIGURE 5 , 7C View FIGURE 7 , 9C View FIGURE 9 ), length/width=0.1–0.2.

Style. Narrow, elongate, greatest length/greatest width=3.4–4.4, length apophysis/greatest length=0.2; apophysis acuminate, apex narrowly rounded, ventrally denticulate, curvate ( Figs 7G View FIGURE 7 , 9G View FIGURE 9 ) or sublinear ( Figs 11L View FIGURE 11 , 13L View FIGURE 13 ), orientation lateroposteriad, angle 38–70°, about half or less as wide as width across preapical lobe; preapical lobe right-angled; anterior medial lobe short in dorsal view; anterior lateral lobe subparallel to base of style ( Figs 9G View FIGURE 9 , 11L View FIGURE 11 , 13L View FIGURE 13 ), about as wide as base or obtusely angled to base ( Fig. 7G View FIGURE 7 ), narrower than base. Apex of apophysis of style inserted either less than halfway ( Figs 7C View FIGURE 7 , 9C View FIGURE 9 ) or three quarters ( Fig. 11C View FIGURE 11 ) length of subgenital plate (halfway to three quarters in C. acapitatus sp. n.).

Connective. Transverse, stem short, wide, arms longer than stem, widely separated, sublinear. Greatest width across arms/width across apex of stem=1.5–2.1. Length arms/length stem=0.7–2.3, variable due to longer or shorter desclerotized margin medially at base of arms, e.g., Figs 7H View FIGURE 7 , 11F View FIGURE 11 , with wide margin; Figs 9N View FIGURE 9 , 11E View FIGURE 11 , 13O, P View FIGURE 13 , with narrower margin. Angle of arms 75–91°. Greatest length/greatest width=0.5–0.6 ( Figs 7H View FIGURE 7 , 9N View FIGURE 9 , 11E, F View FIGURE 11 , 13O, P View FIGURE 13 ).

Aedeagus. Orientation of aedeagus in pygofer with apex of shaft dorsad, vertical, in membranous recess in segment 10. Shaft, dorsal apodeme and preatrium configured as follows:

1. Dorsal apodeme half to two thirds as long as shaft, subparallel to shaft, attached to shaft at atrium, preatrium much shorter than shaft ( Fig. 7D View FIGURE 7 ).

2. Dorsal apodeme about one third as long as shaft, preatrium about as long as shaft ( Fig. 9H, I View FIGURE 9 ).

3. Dorsal apodeme reduced, preatrium as long or longer than shaft ( Figs 11G–K View FIGURE 11 , 13A–F View FIGURE 13 ).

Shaft with teeth in distal half paired or single (possibly due to fusion of contiguous teeth) at anterior or posterior margin, medial paired teeth variable in orientation and origin (anterior, medial or posterior). Base of anterior margin of shaft edentate. Gonopore marginal, wide at apex, anterior subapical margin narrowly incised ( Fig. 13G, I, K View FIGURE 13 ) or not incised ( Figs 7E View FIGURE 7 , 9J View FIGURE 9 ), posterior margin narrowly V-shaped to rounded incision ( Figs 7E View FIGURE 7 , 9J View FIGURE 9 ). Species are distinguished by relative width of apex of shaft in lateral view, compared to medial width ( Figs. 11G–K View FIGURE 11 , 13A–F View FIGURE 13 ), by relative length of shaft to that of the dorsal apodeme ( Figs 7D View FIGURE 7 , 9I View FIGURE 9 ) and by finely denticulate area at anterolateral region of apex of shaft ( Figs 7E View FIGURE 7 , 9K–M View FIGURE 9 ).

Female.

Measurements. As above, female similar to that of male.

Terminalia.

Sternite 7. Transverse, rectangular to isosceles trapezoid (width at posterior margin/width at anterior margin=0.6–0.8, greatest length/greatest width=0.4–0.6) with lateral margins rounded. Three shapes of posterior margins: rounded ( Figs 1G View FIGURE 1 , 2I View FIGURE 2 , 8A View FIGURE 8 , 10A, B View FIGURE 10 , 14D, E View FIGURE 14 ) or with deep ( Fig. 12A–C View FIGURE 12 ) (greatest depth of notch/greatest length laterally=0.61–0.77) or shallow notch ( Fig. 14A–C View FIGURE 14 ) (greatest depth of notch/greatest length laterally=0.75– 0.84), posterior margins laterad of notch rounded.

Valvula 1. Parallel-sided, apex acuminate ( Figs 8I View FIGURE 8 , 10G View FIGURE 10 , 12H View FIGURE 12 , 14F View FIGURE 14 ), sculpture pattern commonly reticulate, striate towards base ( Figs 8K, L View FIGURE 8 , 10K, L, M View FIGURE 10 , 12L, M, N View FIGURE 12 ).

Valvula 2. Serration about in distal half of dorsal margin ( Figs 8H View FIGURE 8 , 10H View FIGURE 10 , 12I View FIGURE 12 , 14J View FIGURE 14 ), narrow, short teeth with trough between successive teeth finely serrate ( Figs 8J View FIGURE 8 , 10N View FIGURE 10 , 12O View FIGURE 12 ).

Valvula 3. 5–22 marginal to submarginal macrosetae at apex, 25–37 µm in length, short setae up to 17 µm ( Figs 8M View FIGURE 8 , 10E, F View FIGURE 10 , 12J, K View FIGURE 12 ). One quarter of apex of valvula 3 protruding beyond pygofer.

Valvifer 1. Rhomboid or rectangular with dorsal and ventral margin somewhat rounded, anterior margin narrowly rounded, posterior margin acute; dorsally fused widely ( Fig. 8B View FIGURE 8 ). Length/width=1.3–1.8 ( Figs 8C, D, E View FIGURE 8 , 10C, D View FIGURE 10 , 12D, E View FIGURE 12 , 14K, L View FIGURE 14 ).

Valvifer 2. Apices narrowly rounded ( Figs 8G View FIGURE 8 , 10I View FIGURE 10 , 12F View FIGURE 12 , 14G View FIGURE 14 ), sculpture pattern at anteroventral margin and submargin reticulate or densely set with microtrichia and few circular structures ( Figs 8F View FIGURE 8 , 10J View FIGURE 10 , 12G View FIGURE 12 , 14H, I View FIGURE 14 ), one specimen with seta. Length/width=2.6–3.0.

Remarks. The combination of external features namely, color pattern of head and tegmina and less so pronotum, and crown angle correspond well within species of Colistra . Other distinguishing features of the genus are given in the diagnosis.

Colistra forms a group with Xhoreus , Proekes and Proekoides , all having some correspondence in size, shape, color pattern and male genitalia. As some specimens appear devoid of the above markings, male genitalia have to be examined. The following similarities and differences are however apparent.

Overall length and body proportions in Colistra bears some similarities with Proekoides with respect to the length from the apex of the crown to the apex of the tegmina, i.e., 3.1–3.5 mm in Proekoides and 2.7–3.4 mm in Colistra . Proekes is much larger (length from apex of crown to apex of abdomen, all specimens, 4.0– 5.1 mm while in the female the ovipositor extends well beyond the apex of the pygofer.

Color pattern of the head, dorsally, includes crown, disc and base tabulated below: View Table

Externally, Colistra also resembles some other Bonaspeiini , e.g., Kimbella Davies, 1988 , Bonaspeia Linnavuori, 1961 (new species in Davies, 1987 and Curvostylus Davies, 1987 . In Kimbella ( Figs 4A, B View FIGURE 4 ), the length from the apex of the crown to the apex of the is tegmina 3.30–3.74 mm, the crown angle is 73–77° and color pattern of head with paired, transverse, deltate marks and small, single, light brown circular marking on the disc. However, in Kimbella the aedeagus has the dorsal apodeme long and preatrium short, and pygofer lobe at posteroventral margin with triangular, serrate process. Tentative differences between Colistra on the one hand and Bonaspeia and Curvostylus on the other, are that the latter two genera lack distinct color pattern on the head, are larger in size, have a long aedeagal shaft, shorter dorsal apodeme and preatrium and different features of the apophysis of the style. With regard to size, length from apex of crown to apex of tegmina (from values in Davies, 1987) in Bonaspeia males 3.46–4.25 mm, females 3.40–3.96 mm, Curvostylus , both sexes, 3.47–4.50 mm and Colistra , both sexes, 2.70–3.20 mm. The apophysis of the style in Bonaspeia could be considered triangular, asymmetric, generally linear, orientation posteriad, about three quarters as wide as width across subapical lobe. The style in Curvostylus has the apophysis digitate, curvate or sublinear, orientation posterolaterad, up to half as wide as width across subapical lobe. Thus, the styles of Colistra and Curvostylus correspond. The aedeagus of most species of Bonaspeia appear to have the dorsal apodeme and preatrium shorter than the shaft and often with the preatrium shorter than the dorsal apodeme, with the exception in B. karensis .The above differentiation of the three genera is tentative due to numerous examined specimens considered new for Bonaspeia and Curvostylus but also probable new genera (personal observation of first author), as well as lack of metrics. Superficial resemblance in marking and color patterns especially on the head are also found in Refrolix trangalensis Theron, 1984 ( Fig. 4D View FIGURE 4 ) and some undetermined Bonaspeiini from the Albany Thicket Biome ( Figs 4C View FIGURE 4 , 5F–I View FIGURE 5 and 6A–H View FIGURE 6 ), and in specimens from the Savanna Biome ( Fig. 6A, B View FIGURE 6 ) with small markings at the apex of the crown. Specimens in Fig. 6F–H View FIGURE 6 ( Fig. 6F View FIGURE 6 , Theewaterskloof, female, Fig. 6G, H View FIGURE 6 , Jonkershoek, male and female respectively) closely resemble the color pattern of the head of Colistra , but are larger (corresponding that of Bonaspeia ( Fig. 5A–C View FIGURE 5 ) and Curvostylus ( Fig. 4E–H View FIGURE 4 )). The male aedeagus of these specimens show some resemblance to the aedeagus in Fig. 13E View FIGURE 13 and with a much longer, single tooth on the preatrium as in Fig. 11K View FIGURE 11 and Fig. 13D View FIGURE 13 .

In male genitalia the pygofer lobe process is present in three of the four genera treated here (absent in Xhoreus gen. n.), but difficult to observe, and is also present in other genera, e.g., Bonaspeia . Most reliable for recognition of genera is the denticulation at the anterior base of the aedeagal shaft, for example, in all species of Colistra , the basal anterior margin of shaft is consistently edentate. Also, the apical, subapical and medial denticulation of the shaft are variable in orientation, position and size and sometimes absent, as discussed under the remarks of each species. In Colistra and Proekes , if the dorsal apodeme is developed the preatrium is reduced and vice versa. In Proekoides , the dorsal apodeme is consistently reduced and the preatrium developed. These features are tabulated as follows:

The connective in Colistra ( Figs7H View FIGURE 7 , 9N View FIGURE 9 , 11E View FIGURE 11 , 13O, P View FIGURE 13 ), Proekoides ( Fig. 16K–O View FIGURE 16 ) and Proekes ( Figs 23H View FIGURE 23 , 26F View FIGURE 26 , 28E View FIGURE 28 , 30J View FIGURE 30 ) is wider than long, with a short, wide stem and widely angled arms. In Xhoreus it least comparable to the former, about as wide as long, narrow stem, but similar angle of arms. Styles of the four genera correspond, with metrics under development.

Key to males of Colistra species

1 Aedeagus with elongate dorsal apodeme; shaft about three quarters longer than dorsal apodeme, preatrium short ( Fig. 7D, E View FIGURE 7 ), apex with anterolateral margin finely denticulate ( Fig. 7D, E View FIGURE 7 )............................... C. parvulus (Linnavuori) View in CoL

1`Aedeagus with short dorsal apodeme, shaft about two thirds longer than dorsal apodeme ( Fig. 9H, I View FIGURE 9 ), or dorsal apodeme reduced ( Figs 11G–K View FIGURE 11 , 13A–F View FIGURE 13 ), apex of shaft variable......................................................... 2

2 Dorsal apodeme of aedeagus short, preatrium elongate, about as long as shaft ( Fig. 9H, I View FIGURE 9 ), shaft with subapex of posterior margin with single tooth and subapex of anterior margin finely denticulate ( Fig. 9K–M View FIGURE 9 )............... C. semialius View in CoL sp. n.

2`Dorsal apodeme of aedeagus reduced, shaft of aedeagus at apex in lateral view of variable width ( Figs 11G–K View FIGURE 11 , 13A–F View FIGURE 13 ).... 3

3 Apex of shaft wide, more than twice as wide as medial width of shaft ( Fig. 11G–K View FIGURE 11 )................. C. bucapitatus View in CoL sp. n.

3`Apex of shaft narrow, up 1.8 times as wide as medial width of shaft ( Fig. 13A–F View FIGURE 13 )................... C. acapitatus View in CoL sp. n.