Australocosmica nana, Criscione, Francesco & Köhler, Frank, 2013

Criscione, Francesco & Köhler, Frank, 2013, Six new species of Australocosmica Köhler, 2011 from the Kimberley islands, Western Australia (Mollusca: Pulmonata: Camaenidae), Zootaxa 3608 (2), pp. 101-115 : 109-111

Australocosmica nana n. sp.

Figs 3 D, 5A–B, 6D

Holotype. Australia, WA, Kimberley, Bonaparte Archipelago, Albert Island: Sta. KC-043, Bonaparte Archipelago, -14.519, 124.929 (coll. V. Kessner, A. Longbottom, 14 July 1988) (WAM S84060 ). Paratypes. Australia, WA, Kimberley, Bonaparte Archipelago, Albert Island: same data as for holotype, 14 wet (WAM S83039 ); 36 dry (WAM S83038 ), 35 dry (FMNH 219165).

Etymology. In reference to its small shell, derived from “ nanus ” (Latin=dwarf), adjective of feminine gender.

Description. Shell ( Figs 3 D; 5A, B). Small, with short spire and sub-globose shape. Whorls with slightly angulate periphery, moderately shouldered underneath suture. Axial ribs moderately pronounced. Colour horn to light brown. Genitalia ( Fig. 6 D). Penis about half as long as anterior part of oviduct (= vagina and free oviduct). Inner penial wall supporting complex sculpture, with series of folds surrounding the opening to vas deferens forming a collar, posteriorly with three corrugated, wide, longitudinal pilasters, anteriorly with numerous, narrow longitudinal pilasters extending into atrium. Vaginal length equivalent to ¾ of penis and free oviduct. Spermoviduct 3.5 times longer than free oviduct and almost twice as long as albumen gland.

Remarks: Shell colour as variable as A. crassicostata . Shell smaller, flatter and with fewer whorls than other congeners. Sculpture similar to that of A. rotunda and with fewer but more pronounced axial ribs than other congeners except A. crassicostata . Inner penial wall sculpture similar to that of A. buffonenis and A. pallida but with more developed transverse folds on lateral pads. This is Solem’s (1991) manuscript species ‘NSP 80’.

Figures

FIGURE 3. Shells of Australocosmica species. A. A. bernoulliensis n. sp. (paratype FMNH 219104). B. A. buffonensis n. sp. (paratype WAM S 83007). C. A. crassicostata n. sp. (paratype FMNH 219181) D. A. nana n. sp. (paratype WAM S 83039). E. A. pallida n. sp. (paratype WAM S 83006). F. A. rotunda n. sp. (paratype WAM S 49237). Scale bar = 10 mm.

FIGURE 6. Penial anatomy of Australocosmica species. A. A. bernoulliensis n. sp. (holotype WAM S 84059). B. A. buffonensis n. sp. (holotype WAM S 84059). C. A. crassicostata n. sp. (holotype WAM S 84059). D. A. nana n. sp. (holotype WAM S 84060). Scale bar = 2 mm.