Hypsiboas benitezi (Rivero, 1961) FAIVOVICH & HADDAD & GARCIA & FROST & CAMPBELL & WHEELER, 2005

FAIVOVICH, JULIÁN, HADDAD, CÉLIO F. B., GARCIA, PAULO C. A., FROST, DARREL R., CAMPBELL, JONATHAN A. & WHEELER, WARD C., 2005, Systematic Review Of The Frog Family Hylidae, With Special Reference To Hylinae: Phylogenetic Analysis And Taxonomic Revision, Bulletin of the American Museum of Natural History 2005 (294), pp. 1-240 : 86-87

Hypsiboas benitezi Group

DIAGNOSIS: This species group is diagnosed by 30 transformations in nuclear and mitochondrial protein and ribosomal genes. See appendix 5 for a complete list of these molecular synapomorphies. A putative synapomorphy of this group is the presence of a flat mental gland in males (Faivovich et al., in prep.) (known instance of homoplasy in Hyla granosa ).

COMMENTS: This new species group includes the clade composed of three species from the Guayana Highlands and three species from the northwestern Amazon Basin. Because one of the Guayanan and one of the Amazonian species are still undescribed, they are not further considered. Two species, Hyla microderma and H. roraima , are a fragment of the former H. geographica group. We are also including tentatively H. hutchinsi and H. rhythmicus , based on their overall similarity with H. benitezi . See appendix 4 for a justification of the inclusion of Hyla pulidoi (Rivero, 1968) in this species group.

CONTENTS: Seven species. Hypsiboas benitezi (Rivero, 1961) , new comb.; Hypsiboas hutchinsi (Pyburn and Hall, 1984) , new comb.; Hypsiboas lemai (Rivero, 1971) , new comb.; Hypsiboas microderma (Pyburn, 1977) , new comb.; Hypsiboas pulidoi (Rivero, 1968) , new comb.; Hypsiboas rhythmicus (Señaris and Ayarzagüena. 2002) , new comb.; Hypsiboas roraima (Duellman and Hoogmoed, 1992) , new comb.