Sagola ignota Broun, 1921

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 93-94

Sagola ignota Broun, 1921

( Figs. 52e, 52k, 52n, 52q–r , 53 )

Sagola ignota Broun, 1921a: 495 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.

Type Material. Holotype. NEW ZEALAND: Otago Lakes : 1♂ ( BMNH), aedeagus dissected and mounted in balsam on a clear plastic card, “4004. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Routeburn. 16.2.1914.” [white label, handwritten]; “ Sagola ignota . ♂ ” [white label, handwritten].

Additional Material (n = 15: 8 males; 7 females). NEW ZEALAND: Dunedin: 1♂, Woodside Glen Outram , 30 XI 2005, J. Nunn, in moss and lichen on tree trunk ; Fiordland: 2♂♂ (1♂, slidemounted), Borland Saddle , c 1050m, 16 XII 1998, in moss on Nothofagus bole, J. Nunn ; 3♀♀, Borland Saddle , 24 I 2008, J. Nunn, sifted moss ; 2♀♀, W. Olivine Ra, Simonin Pass , 23 I 1975, G.W. Ramsay, litter 75/35 ; 1♀, Outer Gilbert 2, Breaksea Sound , Nov 1985, on Kiekie ; 1♂, Fiordland NP, S Borland V Biv , 730m, 2–6 II 1982, J.W. Early, Malaise trap at Nothofagus forest edge ; 1♂, Gertude V, Milford Rd , 23 I 2008, J. Nunn, in moss on beech tree ; 1♂, Barrier Ra, Little Red Hill, Mt. Allison , 4 II 1972, J.S. Dugdale, little 75/52 ; Stewart Island: 1♂, ne Big S. Cape I, 122m, 18 XI 1968, G. Kuschel, litter 68/191 ; 1♀, Island Hill Homestead , 2–8 II 1991, C.J. Vink, J.W. Early, yellow pan trap in bush ; Southland: 1♂, 1km S Longwood Trig, 700m, Longwood Ra, 1 II 1976, L.L. Deitz, mats 76/12 .

Diagnosis. This species can be distinguished from the other species of the furcata speciesgroup by the following combination of characters:antennomeres 4–7 subquadrate; frontal sulcus inverted V-shaped; eye prominent, as long as temple; male abdominal ventrites V–VI flattened roundly, surrounded by dense setae; shape of genitalia unique to species.

Redescription. Length 1.9–2.2 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 52e ). Head: Bluntly triangular, widest across eyes ( Fig. 52q ). Ventral surface of male head with inverted triangular process bearing dense setae behind mouthparts ( Fig. 52r ). Antennomere 1 approximately 1.5 times longer than wide, 2–10 subquadrate ( Fig. 52n ). Frontal sulcus inverted V-shaped ( Fig. 52q ). Anterior frontal fovea rectangular, covered by frontal rostrum ( Fig. 52q ), posterior frontal fovea absent ( Fig. 52q ). Eye prominent, approximately as long as temple ( Fig. 52q ). Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular ( Fig. 52e ). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Male abdominal ventrites V–VI roundly flattened on middle with dense setae. Aedeagus: Median lobe of genitalia weakly curved and long with V-shaped depression ( Fig. 52k ). Phallobase of median lobe symmetrical and rounded ( Fig. 52k ). Parameres symmetrical and as long as median lobe, with setae apically ( Fig. 52k ).

Distribution. Dunedin, Fiordland, Stewart Island, Southland ( Fig. 53 : white circles).

Habitat. Specimens of this species were collected using Malaise and yellow pan traps or by sifting leaf, moss, and leaf litter in Nothofagus forests.

Figures

Fig. 52. Sagola species in the furcata species-group. Habitus, dorsal views: a) S. furcata, b) S. punctulata, c) S. pecki, d) S. muirae, e) S. ignota, f) S. lewisensis. Aedeagi, dorsal views: g) S. furcata, h) S. punctulata, i) S. pecki, j) S. muirae, k) S. ignota, l) S. lewisensis. m) S. lewisensis, aedeagus, lateral view, n) S. ignota, antenna. S. punctulata, male head: o) Dorsal surface (arrow indicates inverted Y-shaped frontal sulcus), p) Ventral surface. S. ignota, male head: q) Dorsal surface (arrow indicates inverted V-shaped frontal sulcus), r) Ventral surface. Scale bars a–f = 1 mm, g–r = 0.1 mm.

Fig. 53. Known collection localities of New Zealand Sagola species. S. furcata: black circles; S. punctulata: triangles; S. pecki: black square; S. muirae: stars; S. ignota: white circles; S. lewisensis: white squares.