Sagola tenebrica Broun, 1921

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 74

Sagola tenebrica Broun, 1921

( Figs. 41b, 41d , 42 )

Sagola tenebrica Broun, 1921a: 487 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244.

Type Material. Holotype. NEW ZEALAND: Mid Canterbury: 1♀ ( BMNH), glued on rectangular card, “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Moa Basin. 20.10.1913.” [white label, handwritten]; “ Sagola ♀ tenebrica.” [white label, handwritten].

Additional Material (n = 19; 11 males, 8 females). NEW ZEALAND: Fiordland : 4♂♂ 6♀♀, Borland Saddle , 24 I 2008, sifted moss, J, Nunn ; 1♀, Borland Saddle , 1050m, 16 XII 1998, in moss on Nothofagus bole, J. Nunn ; 1♂, Breaksea So , 12 III 1983, C.F. Butcher ; 1♂, Secretary I, Mt. Grono , 853m, 27 XI 1981, C.F. Butcher, mats, moss, tussock 91/186 ; 1♂, Fiordland NP, Milford Sound rd, Gertrude Saddle car park area, 800m, 44″46.13′S, 168″00.16′E, 9 XII 2005, very mossy Nothofagus menziesii forest, FMHD2005-088 , recent flood debris floating in stream, J. Nunn, A. Newton, ANMT site 1169; Mid Canterbury: 1♂, Arthur Pass, Dobson Nature Walk , 8 II 1982, C.F. Butcher, sifted litter 82/26 ; 1♂, Moa Basin , A.E. Brookes Collection, T. Broun Collection ; Mackenzie: 1♂, Mt. Burns , 1700m, 13 I 1970, J.I. Townsend, Mats 70/101 ; Nelson: 1♂, Mt. St. Arnaud , 1900m, 1 VII 1973, J.C. Watt, litter 73/109 ; Westland: 1♂, Fox G 1200m, Chancellor Hut , 15 XII 1984, J.W. Early, P Syrett, Subalpine herb field tussock litter LCNZ 84 /8 .

Diagnosis. This species can be distinguished from the other species of the arboricola speciesgroup by the following combination of characters: larger body length, 2.5–3.0 mm; ventral surface of male head with blunt process bearing dense setae behind mouthparts; male antennomeres 2–10 subquadrate and weakly enlarged; shape of genitalia unique to species.

Redescription. Length 2.5–3.0 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 41b ). Head: Bluntly triangular, widest across eyes ( Fig. 41b ). Ventral surface of male head with blunt process bearing dense setae behind mouthparts. Male antennomere 1 approximately 1.5 times longer than wide, 2–10 subquadrate and weakly enlarged. Female antennomeres 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate. Frontal sulcus deep, reaching one-third length of eye. Anterior frontal fovea oval, posterior frontal fovea round. Eye large and prominent, approximately as long as temple. Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular ( Fig. 41b ). Mesoand metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle, less developed in female. Aedeagus: Median lobe triangular, dorsal process crescent-shaped ( Fig. 41d ). Phallobase of median lobe symmetrical and rectangular ( Fig. 41d ). Parameres symmetrical and rectangular with sparse setae anteriorly and posteriorly ( Fig. 41d ).

Type Locality. Moa Basin , Mid Canterbury, New Zealand .

Distribution. Fiordland, Mid Canterbury, Mackenzie, Nelson, Westland ( Fig. 42 : triangles).

Habitat. Most specimens of this species were collected by sifting mossy leaf or log litter in Nothofagus forests. Many specimens were collected at high elevations.

Figures

Fig. 41. Sagola species in the arboricola species-group, Habitus, dorsal views: a) S. arboricola, b) S. tenebrica. Aedeagi, dorsal views: c) S. arboricola, d) S. tenebrica. S. arboricola, male head: e) Dorsal surface, f) Ventral surface. Scale bars a–b = 1 mm, c–f = 0.1 mm.

Fig. 42. Known collection localities of New Zealand Sagola species. S. arboricola: black circles; S. tenebrica: triangles.