Jurochlus adustus, Lukashevich, Elena D. & Przhiboro, Andrey A., 2012

Lukashevich, Elena D. & Przhiboro, Andrey A., 2012, Pupae of Mesozoic Jurochlus Kalugina, 1985 (Diptera: Chironomidae), with description of four new species, Zootaxa 3478, pp. 434-452 : 446

Jurochlus adustus sp. nov.

( Figs 3 I, J, 8B, C)

Type material. Holotype PIN 3965/3209 (female pupal abdominal segments III–IX). Mongolia, East Gobi Aymag, Khutel-Khara (=Hara Hutul), lower Tsagantsab Formation, Lowermost Cretaceous (Uppermost Jurassic not excluded).

Etymology. From the Latin adustus (swarthy), in reference to dark pattern.

Description. Female. Anterior margins of tergites appearing as wide slightly sinuate sclerotized stripes; posterior margins of all segments with a thin line, without traces of ridge. Posterolateral margins of tergites, as a rule, slightly projecting, rounded or bluntly tapered. Tergites uniformly moderately darkened except for lighter lateral margins; posterior margins slightly more darkened. LS setae and their attachment places not visible. Tergites III–VII with traces of short setae scattered over surface, most clear on segment V. Segments III–V posterior to hind margin of tergite (probably intersegmental membrane) with fine, more or less longitudinal (fan-shaped) striation (possibly, traces of muscles); some of such lines extending to posterior or anterior margins of tergites.

Segment IX. Outer margins of anal lobes almost straight in apical two-thirds, slightly concave near apex, possibly indistinctly serrate in middle parts (no distinct teeth visible). Inner margins slightly sinuate. Anal lobes with strongly impressed stout spurs. Genital sac tapering, apically more or less truncate. Genital plate IX appearing as transverse stripe with margins slightly impressed.

Measurements. Combined length of abdominal segments IV–IX, 5500; entire body length (extrapolated), 9500–10500. Abdominal segments length: IV–VII, 800; VIII, 870; width: III–VII, 2150–2200; VIII, 1920. Abdominal segment IX: length, 1250; width at base, 1500; distance between tips of anal lobe spurs, 500.

Figures

FIGURE 3. SEM images of Jurochlus pupae from the Mesozoic of Mongolia: A – E, J. trivittatus sp. nov.: A – C, holotype PIN 4270 / 2581, female: A, total view (SE), B, outer margin of anal lobe (BSE), C, serration of outer margin of anal lobe (BSE); D, E, paratype PIN 4270 / 2540, male: D, apex of abdomen (SE), E, outer margin of anal lobe (SE); F – H, J. limbatus sp. nov., holotype PIN 4270 / 2584, female: F, total view (BSE), G, outer margin of anal lobe (BSE), H, lateral margin of segment VIII (BSE); I, J, J. adustus sp. nov., holotype PIN 3965 / 3209, female: I, apex of abdomen (BSE), J, segments III, IV, V (BSE).